Aix-en-Provence (MNHN Paris Coll) (Oligocene of France)

Where: Bouches-Du-Rhone, France (43.5° N, 5.5° E: paleocoordinates 43.3° N, 2.8° E)

• coordinate estimated from map

• local area-level geographic resolution

When: Niveau du gypse d'Aix Formation, Chattian (28.1 - 23.0 Ma)

• "The ‘insect formation’ is 2.5 m thick, divided in two parts that correspond to two very different environmental conditions. The first (lower) episode is remarkable by its lateral monotony of facies on a distance of 6 km at least, in the outcrops observed on the northwest of Aix-en-Provence. This lower episode is distinguished from the rest of the ‘insect formation’ and from the whole formation of Aix-en-Provence by the presence of a very numerous thin limestone laminae. The second (upper) episode is remarkable by the presence of many silici- fied laminae. The fauna in this level is also very well preserved"….This collection consists of unspecified layers

Environment/lithology: lacustrine - large; lithified limestone

• This lacustrine formation was deposited in a very large but shallow-water lake, with variable salinity and near the marine coast (Nury 1990, Nury & Thomassin 1994). Irregular basins formed during extension of the European platform in the Oligocene resulting from the formation of the West-European rift system.
• The deposit presents an alternation of limestone and marls with gypsum.

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Reposited in the MNHN

Primary reference: F. Meunier. 1915. Nouvelles recherches sur quelques insectes des Plâtrières d'Aix en Provence. Verhandelingen der Koninklijke Akademie van Wetenschappen te Amsterdam 18(5):1-17 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 122501: authorized by Matthew Clapham, entered by Jered Karr on 31.12.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera - Lygaeidae
Diniella fasciata13 Heer 1853 milkweed bug
Coll Museum Paris Am 147-151
Heterogaster antiquus13 Heer 1853 milkweed bug
Coll Museum Paris Am 127
Heterogaster pumilio13 Heer 1853 milkweed bug
Coll Museum Paris Am 140-142
Aphanus pulchellus13 Heer 1853 milkweed bug
Coll Museum Paris Am 143-146, 138
Aphanus dilatatus n. sp.13 Theobald 1937 milkweed bug
Coll Museum Paris Am 132
Aphanus murchisonae13 Heer 1853 milkweed bug
Coll Museum Paris Am 122-126, 100-121
 Hemiptera - Berytidae
? Neides oligocenicus n. sp.5 Nel 1992 stilt bug
MNHN IPM-R.10372, 10373, 10374, 10375
 Hemiptera - Cydnidae
Cydnus meunieri13 Theobald 1937 burrowing bug
Coll Museum Paris Am 139
 Hemiptera - Hydrometridae
Protobacillometra oligocenica n. gen. n. sp.9 Nel and Paicheler 1993 marsh treader
MNHN LP-R.07893
 Blattodea -
Paleotermopsis oligocenicus n. gen. n. sp.8 Nel and Paicheler 1993 termite
MNHN IPM-R.54190
Ulmeriella aquisextana n. sp.4 Nel 1986 termite
MNHN IPM-R.07443 (Nel no. 1358); IPM-R.54242 in Nel & Paicheler 1993
 Blattodea - Mastotermitidae
Mastotermes sp.8 Froggatt 1896 termite
MNHN IPM-R.54195, 54324
Mastotermes gallica n. sp.4 Nel 1986 termite
MNHN IPM-R.07441 (Nel no. 1)
Blattotermes massiliensis n. sp.4 Nel 1986 termite
MNHN IPM-R.07442 (Nel no. 1380)
 Dermaptera - Forficulidae
Forficula "sp. A"6 Linnaeus 1758 common earwig
MNHN-LP-R.10401
 Orthoptera - Gryllidae
Gryllidae indet.14 Laicharting 1781 true crickets
in the collection of Oustalet
 Hymenoptera - Pompilidae
"Pompilus fasciatus n. sp." = Pompilites fasciatus13, "Pompilus coquandi n. sp." = Pompilinites coquandi13
"Pompilus fasciatus n. sp." = Pompilites fasciatus13 Theobald 1937 spider wasp
Coll Museum Paris Am 11
"Pompilus coquandi n. sp." = Pompilinites coquandi13 Theobald 1937 spider wasp
Coll Museum Paris Am 1
 Hymenoptera - Mutillidae
Methoca sp.13 Latreille velvet ant
Coll Museum Paris Am 3
 Hymenoptera - Formicidae
Lasius epicentrus n. sp.13, "Formica minutula" = Lasius occultatus13, Formica sp.13, Formica sepulta n. sp.13, Formica latinodosa n. sp.13, "Camponotus saussurei n. sp." = Camponotus theobaldi13, Camponotus longiventris n. sp.13, Formica oculata13, Dolichoderus coquandi n. sp.13, "Aphaenogaster maculata" = Paraphaenogaster maculata13, "Brachyponera dubia" = Pachycondyla dubia13
Lasius epicentrus n. sp.13 Theobald 1937 ant
Coll Museum Paris Am 7
"Formica minutula" = Lasius occultatus13 Heer 1849 ant
Coll Museum Paris Am 15
Formica sp.13 Linnaeus 1758 ant
Coll Museum Paris Am 6
Formica sepulta n. sp.13 Theobald 1937 ant
Coll Museum Paris Am 5
Formica latinodosa n. sp.13 Theobald 1937 ant
Coll Museum Paris Am 9
"Camponotus saussurei n. sp." = Camponotus theobaldi13 Özdikmen 2010 ant
Coll Museum Paris Am 4
Camponotus longiventris n. sp.13 Theobald 1937 ant
Coll Museum Paris Am 2, 18
Formica oculata13 Heer 1849 ant
Coll Museum Paris Am 19
Dolichoderus coquandi n. sp.13 Theobald 1937 ant
Coll Museum Paris Am 22
"Aphaenogaster maculata" = Paraphaenogaster maculata13 Theobald 1937 ant
Coll Museum Paris Am 10
"Brachyponera dubia" = Pachycondyla dubia13 Theobald 1937 ant
Coll Museum Paris Am 20
 Hymenoptera - Apidae
Apis "sp. B"7, Apis "sp. C"7, "Apis aquisextusensis n. sp." = Apis (Synapis) henshawi7
Apis "sp. B"7 Linnaeus 1758 honey bee
MNHN-LP-R.10429
Apis "sp. C"7 Linnaeus 1758 honey bee
MNHN-LP-R.10426
"Apis aquisextusensis n. sp." = Apis (Synapis) henshawi7 Cockerell 1907 Henshaw's honey bee
MNHN-LP-R.08381, 08382, 10421
 Hymenoptera - Braconidae
"Diospilus verus n. sp." = Taphaeus verus13
"Diospilus verus n. sp." = Taphaeus verus13 Theobald 1937 wasp
Coll Museum Paris Am 23
 Hymenoptera - Ichneumonidae
Exetastes postornata n. sp.13 Theobald 1937 ichneumon wasp
Coll Museum Paris Am 16
? Demophorus fumipennis n. sp.13 Theobald 1937 ichneumon wasp
Coll Museum Paris Am 14
Ophion annulatus n. sp.13 Theobald 1937 ichneumon wasp
Coll Museum Paris Am 13
Nemeritis longicornis n. sp.13 Theobald 1937 ichneumon wasp
Coll Museum Paris Am 12
? Pimpla saussurei13 Heer 1856 ichneumon wasp
Coll Museum Paris 17
Pimpla antiquus13 Saussure 1852 ichneumon wasp
Coll Museum Paris 21
Promethes tilloyi n. sp.13 Theobald 1937 ichneumon wasp
Coll Museum Paris Am 8
 Coleoptera - Staphylinidae
Staphylinus priscus11, Staphylinus calvus11, Staphylinus aquisextanus n. sp.11, Staphylinus prodromus11, "Staphylinus (Ocypus) provincialis n. sp." = Ocypus provincialis11, "Erinnys deleta n. sp." = Lithoplanes deleta11, Stenus gypsi n. sp.11
Staphylinus priscus11 Oustalet 1874 rove beetle
Staphylinus calvus11 Oustalet 1874 rove beetle
Staphylinus aquisextanus n. sp.11 Oustalet 1874 rove beetle
Staphylinus prodromus11 Oustalet 1874 rove beetle
"Staphylinus (Ocypus) provincialis n. sp." = Ocypus provincialis11 Oustalet 1874 rove beetle
"Erinnys deleta n. sp." = Lithoplanes deleta11 Oustalet 1874 rove beetle
Stenus gypsi n. sp.11 Oustalet 1874 rove beetle
 Coleoptera - Chrysomelidae
Cassida blancheti11, Chrysomela matheroni11, Gonioctena curtisii n. sp.11, "Chrysomela matrona n. sp." = Plagiodera lyelliana11
Cassida blancheti11 Heer 1856 tortoise beetle
Chrysomela matheroni11 Oustalet 1874 leaf beetle
Gonioctena curtisii n. sp.11 Oustalet 1874 leaf beetle
"Chrysomela matrona n. sp." = Plagiodera lyelliana11 Heer 1856 leaf beetle
 Coleoptera - Curculionidae
Hipporhinus heerii11, Hipporhinus heeri13, "Sitones margarum" = Protainophthalmus margarus11, "Brachyderes aquisextanus" = Protainophthalmus asperulus11, "Brachyderes longipes" = Protainophthalmus asperulus11, "Cleonus marcelli" = Protainophthalmus asperulus11, "Hipporhinus reynesii" = Rhytidoderes sexsulcatus11, Cleonus inflexus11
Hipporhinus heerii11 Germar 1849 snout beetle
Hipporhinus heeri13 Germar 1849 snout beetle
"Sitones margarum" = Protainophthalmus margarus11 Germar 1849 snout beetle
"Brachyderes aquisextanus" = Protainophthalmus asperulus11 Heer 1856 snout beetle
"Brachyderes longipes" = Protainophthalmus asperulus11 Heer 1856 snout beetle
"Cleonus marcelli" = Protainophthalmus asperulus11 Heer 1856 snout beetle
"Hipporhinus reynesii" = Rhytidoderes sexsulcatus11 Heer 1856 snout beetle
Cleonus inflexus11 Oustalet 1874 snout beetle
 Coleoptera - Geotrupidae
Geotrupes atavus n. sp.11 Oustalet 1874 earth-boring dung beetle
 Coleoptera - Carabidae
Polystichus hopei n. sp.11 Oustalet 1874 ground beetle
 Diptera -
? Anthomyia pulcherrima13 Meunier 1915 fly
Coll Museum Paris Am 26, 37
 Diptera - Bombyliidae
Exoprosopa romani13 Theobald 1937 bee fly
 Diptera - Syrphidae
"Chilosia oligocenica n. sp." = Cheilosia oligocenica13
"Chilosia oligocenica n. sp." = Cheilosia oligocenica13 Theobald 1937 hoverfly
Coll Museum Paris Am 24
 Diptera - Sciomyzidae
Tetanocera alireticulata n. sp.13 Theobald 1937 snail-killing fly
Coll Museum Paris Am 25
 Diptera - Tabanidae
Chrysops sp.13 Meigen 1803 deer fly
Coll Museum Paris Am 28(29)
 Diptera - Bibionidae
Penthetria nervisinuata12, "Plecia cf. lugens" = Penthetria lugens13, "Plecia livida" = Plecia livida13, "Plecia brevipennis" = Plecia livida13, "Plecia splendida" = Plecia foersteri13, "Plecia bucklandi" = Plecia bucklandi13, "Plecia elegans" = Plecia bucklandi13, "Plecia dilatativentris" = Plecia bucklandi13, "Plecia gracilis" = Plecia bucklandi13, "Plecia guinieri" = Plecia lygaeoides13, Bibiodes provincialis n. sp.12, Bibio sp.13, Bibio spectabilis13, Bibio aquaesextiae n. sp.12, Bibio curtisi13, Bibio disjectus13, "Bibio curtisii" = Bibio curtisii12, "Bibio serresi" = Bibio curtisii13, "Bibio romani" = Bibio curtisii13, Dilophus sp.12
Penthetria nervisinuata12 Theobald 1937 march fly
MNHN Aix 332, 428, 645, 752, 779, 799, 884, 890, 926, 965, 974, 1013, 1061,1085, 1332, 1337, 1347, 1404, P26
"Plecia cf. lugens" = Penthetria lugens13 Oustalet 1870 march fly
Coll Museum Paris Am 39, 41
"Plecia livida" = Plecia livida13 Heer 1849 love bug
Coll Museum Paris Am 51-56, 58, 60
"Plecia brevipennis" = Plecia livida13 Heer 1849 love bug
Coll Museum Paris Am 65
"Plecia splendida" = Plecia foersteri13 Theobald 1937 love bug
Coll Museum Paris Am 63
"Plecia bucklandi" = Plecia bucklandi13 Heer 1849 love bug
Coll Museum Paris Am 50, 19
"Plecia elegans" = Plecia bucklandi13 Heer 1849 love bug
Coll Museum Paris Am 43-46, 59
"Plecia dilatativentris" = Plecia bucklandi13 Heer 1849 love bug
Coll Museum Paris Am 42, 48
"Plecia gracilis" = Plecia bucklandi13 Heer 1849 love bug
Coll Museum Paris Am 57
"Plecia guinieri" = Plecia lygaeoides13 Heer 1849 love bug
Coll Museum Paris Am 64
Bibiodes provincialis n. sp.12 Skartveit and Nel 2017 march fly
MNHN Aix 1079, 23, 62, 543, 676, 959
Bibio sp.13 Geoffroy 1762 march fly
Coll Museum Paris Am 21
Bibio spectabilis13 Theobald 1937 march fly
Coll Museum Paris Am 78; 15 in Skarveti & Nel 2017 (Aix 6, 10, 11, 52, 107, 161, 309, 845, 1090, 1299, 1307, 1377, 1446, 1447, P25)
Bibio aquaesextiae n. sp.12 Skartveit and Nel 2017 march fly
MNHN Aix 152, 309, 416,421, 532, 610, 1010, 1014, 1294, 1436, 1452
Bibio curtisi13 Heer 1856 march fly
Coll Museum Paris Am 77
Bibio disjectus13 Theobald 1937 march fly
Coll Museum Paris Am 75
"Bibio curtisii" = Bibio curtisii12 Heer 1856 march fly
MNHN Aix 42, 54, 584, 873, 950, B.24431
"Bibio serresi" = Bibio curtisii13 Heer 1856 march fly
Coll Museum Paris Am 67-73
"Bibio romani" = Bibio curtisii13 Heer 1856 march fly
Coll Museum Paris Am 74
Dilophus sp.12 Meigen 1803 march fly
MNHN Aix 18, 31,34, 491, 797, 1079
 Diptera - Mycetophilidae
Mycetophila confusa13 Theobald 1937 fungus gnat
Coll Museum Paris Am 30
Mycetophila cf. morio13 Heer 1856 fungus gnat
Coll Museum Paris Am 34-36
Mycomyia curvithoracis n. sp.13 Theobald 1937 fungus gnat
Coll Museum Paris Am 27
"Mycetophila meigeniana" = Gnoriste meigeniana13 Heer 1856 fungus gnat
Coll Museum Paris Am 32
 Diptera - Sciaridae
Sciara oblonga13 Theobald 1937 dark-winged fungus gnat
Coll Museum Paris Am 31, 33
 Lepidoptera - Pterophoridae
"Pterophorus oligocenicus n. sp." = Merrifieldia oligocenicus2
"Pterophorus oligocenicus n. sp." = Merrifieldia oligocenicus2 Bigot et al. 1986 moth
Oustalet collection
 Odonata - Platycnemididae
Platycnemididae indet.10 Jacobson and Bianchi 1905 white-legged damselfly
MNHN IPM-R07704
 Odonata - Coenagrionidae
Coenagrionidae "indet. C"10 Kirby 1890 narrow-winged damselfly
MNHN IPM-R07460
Coenagrionidae "indet. D"10 Kirby 1890 narrow-winged damselfly
MNHN IPM-R07706
Argiinae "indet. A"10 Tillyard 1917 narrow-winged damselfly
MNHN IPM-R07449
Argiinae "indet. B"10 Tillyard 1917 narrow-winged damselfly
MNHN IPM-R07705
 Odonata - Libellulidae
Libellulidae "larva"13 Rambur 1842 common skimmer
Am 80-87
Arachnida
 Araneae - Salticidae
Attoides eresiformis n. gen. n. sp.3
Attoides eresiformis n. gen. n. sp.3 Brongniart 1877 spider
In "Le Muséum", sent by M. Gaudry
 Araneae - Gnaphosidae
Drassodes sextii n. sp.1 Berland 1939 spider