Bruay-la-Bussière, Terril 5bis, Pas-de-Calais (Carboniferous of France)

Where: France (50.5° N, 2.5° E: paleocoordinates 2.3° S, 11.0° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Westphalian A (318.7 - 316.9 Ma)

• "Terril no. 5", black Carboniferous shales, "Faisceau de Modeste", "Veine Maroc" Early Langsettian (Early Westphalian A, Bashkirian Stage).

Environment/lithology: terrestrial; lithified shale

• Variscan foreland basin

Size class: macrofossils

Preservation: mold/impression

Collected by Patrick Roques

Primary reference: A. Nel, P. Roques, P. Nel, J. Prokop, and J. S. Steyer. 2007. The earliest holometabolous insect from the Carboniferous: a "crucial" innovation with delayed success (Insecta Protomeropina Protomeropidae). Annales de la Société Entomologique de France 43(3):349-355 [M. Clapham/M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 122506: authorized by Matthew Clapham, entered by Matthew Clapham on 31.12.2011

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Protomeroptera - Protomeropidae
Westphalomerope maryvonneae n. gen. n. sp. Nel et al. 2007 winged insect
 Caloneurodea -
Oudardgramma bruayensis n. gen. n. sp.
Oudardgramma bruayensis n. gen. n. sp. Nel and Roques 2021 winged insect
MNHN.F.A71355