Hadjoula (Day collection, AMNH) (Cretaceous of Lebanon)

Also known as Hjoula, Hgula, Hajula, Hadjula, Djoula

Where: Lebanon (34.1° N, 35.7° E: paleocoordinates 12.9° N, 30.4° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Late/Upper Cenomanian (99.6 - 93.5 Ma)

• Hjoûla Cretaceous limestone

• group of beds-level stratigraphic resolution

Environment/lithology: lithified limestone

• The limestone sedimentation including fossils is not flat: in fact, bending and fractures, with frequent shifting, are common events within very small areas at the Hgula quarries. This non-homogeneous limestone configuration does not permit the recovery of large slabs of limestone. On the other hand, the Hgula limestone is characterized by an impressive richness: dozen of fossil fishes almost completely cover each limestone surface, with frequent overlapping of individual specimens. The most frequent fossils are undoubtedly fishes, but shrimps are also very common.

Size class: macrofossils

Collected by Alfred Ely Day in 1901; reposited in the AMNH

Primary reference: O. P. Hay. 1903. On a collection of upper Cretaceous fishes from Mount Lebanon, Syria, with descriptions of four new genera and nineteen new species. Bulletin of the American Museum of Natural History 19:395-452 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 221229: authorized by Matthew Clapham, entered by Matthew Clapham on 17.07.2021

Creative Commons license: CC BY (attribution)

Taxonomic list

Chondrichthyes
 Rajiformes - Rhinobatidae
Rhinobatus eretes n. sp., "Raja whitfieldi n. sp." = Rhinobatos whitfieldi
Rhinobatus eretes n. sp. Hay 1903 guitarfish
AMNH 4500
"Raja whitfieldi n. sp." = Rhinobatos whitfieldi Hay 1903 guitarfish
AMNH 4504a, b, c
 Rajiformes - Sclerorhynchidae
"Sclerorhynchus hiram n. sp." = Libanopristis hiram, Sclerorhynchus solomonis n. sp., Sclerorhynchus sentus n. sp.
"Sclerorhynchus hiram n. sp." = Libanopristis hiram Hay 1903 ray
AMNH 4501
Sclerorhynchus solomonis n. sp. Hay 1903 ray
AMNH 4503
Sclerorhynchus sentus n. sp. Hay 1903 ray
AMNH 4504
Actinopteri
 Ichthyodectiformes - Cladocyclidae
Eubiodectes libanicus Pictet and Humbert 1866
AMNH 4506b
 Aulopiformes - Enchodontidae
Enchodus ? marchesettii Gorjanović-Kramberger 1895
AMNH 4527b, c
 Aulopiformes - Aulopidae
Nematonotus longispinus Davis 1887
AMNH 4510a-f
 Aulopiformes - Ichthyotringidae
"Rhinellus delicatus n. sp." = Ichthyotringa delicata
"Rhinellus delicatus n. sp." = Ichthyotringa delicata Hay 1903
AMNH 4530
 Aulopiformes - Dercetidae
"Leptotrachelus serpentinus n. sp." = Benthesikyme serpentina
"Leptotrachelus serpentinus n. sp." = Benthesikyme serpentina Hay 1903
AMNH 4511a
 Aulopiformes -
Acrognathus dodgei n. sp. Hay 1903
AMNH 4520a, b
 Myctophiformes - Sardinioididae
"Osmeroides pontivagus n. sp." = Sardinioides minimus
"Osmeroides pontivagus n. sp." = Sardinioides minimus Agassiz 1839
AMNH 4524a, c, e, f
 Polymixiiformes - Pycnosteroididae
"Pycnosterinx levispinosus n. sp." = Pycnosteroides levispinosus, "Aipichthys formosus n. sp." = Pycnosteroides levispinosus
"Pycnosterinx levispinosus n. sp." = Pycnosteroides levispinosus Hay 1903 beardfish
AMNH 4528
"Aipichthys formosus n. sp." = Pycnosteroides levispinosus Hay 1903 beardfish
AMNH 4519
 Ctenothrissiformes - Ctenothrissidae
Ctenothrissa signifer n. sp. Hay 1903
AMNH 4521a, b
 Elopiformes - Megalopidae
"Microcoelia dayi n. sp." = Sedenhorstia dayi
"Microcoelia dayi n. sp." = Sedenhorstia dayi Hay 1903
AMNH 4525a, b, c, d
 Anguilliformes -
"Urenchelys germanus n. sp." = Hayenchelys germanus
"Urenchelys germanus n. sp." = Hayenchelys germanus Hay 1903 eel
AMNH 4515a, b, c, e
 Anguilliformes - Anguillavidae
"Anguillavus quadripinnis n. sp." = Anguillavus quadripinnis, "Anguillavus bathshebae" = Anguillavus quadripinnis
"Anguillavus quadripinnis n. sp." = Anguillavus quadripinnis Hay 1903 eel
AMNH 4512
"Anguillavus bathshebae" = Anguillavus quadripinnis Hay 1903 eel
AMNH 4513b, c
 Pycnodontiformes - Pycnodontidae
Coccodus insignis n. sp. Hay 1903
AMNH 4516a, b, c, d, e, f