El Horcajo (Cretaceous of Spain)

Also known as Trevijano, TRE, TREN

Where: La Rioja, Spain (42.3° N, 2.4° W: paleocoordinates 33.6° N, 8.1° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: El Horcajo Unit Formation (Enciso Group), Late/Upper Berriasian (145.0 - 140.2 Ma)

• "El Horcajo Unit" overlies the Leza Formation; some authors place it within the Oncala Group

• bed-level stratigraphic resolution

Environment/lithology: lacustrine delta plain; gray marl and red claystone

• "lacustrine-palustrine environment"
• "a grey marls level that is included in a red clay-grey marl interval"

Size classes: macrofossils, mesofossils, microfossils

Collected by A. Fernández in the 1990s

Collection methods: bulk, surface (in situ), sieve,

Primary reference: M. Moreno-Azanza, J. M. Gasca, I. Díaz-Martínez, A. Fernández, J. I. Canudo and F. Pérez-Lorente. 2013. Ootaxa assemblage from El Horcajo site (upper Berriasian, Cameros Basin) in Trevijano (La Rioja, Spain). In F. Torcida Fernández-Baldor & P. Huerta (ed.), VI Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. Libro de Resúmenes 93-95 [M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 224944: authorized by Matthew Carrano, entered by Matthew Carrano on 13.04.2022

Creative Commons license: CC BY (attribution)

Taxonomic list

 Charales - Clavatoraceae
Atopochara trivolvis Peck 1938
A. t. horrida, A. t. micranda, A. t. ancora
Echinoidea
 Echinoneoida - Conulidae
Globator maillardii sea urchin
G. m. incrassatus, G. m. nurrensis
Reptilia
 Testudinata - Testudoolithidae
? Testudoolithidae indet.1 Hirsch 1996 turtle
 Ornithischia -
Ornithopoda indet. Marsh 1881 ornithopod
vertebra
 Ornithischia - Hypsilophodontidae
""Hypsilophontidae" indet." = Hypsilophodontidae
""Hypsilophontidae" indet." = Hypsilophodontidae Dollo 1882 hypsilophodontid
teeth
 Ornithischia - Spheroolithidae
Guegoolithus turolensis1 Amo Sanjuán et al. 2000 ornithopod
  - Prismatoolithidae
Prismatoolithidae indet. Hirsch 1994 dinosaur
 Theropoda - Spinosauridae
Spinosauridae indet.2 Stromer 1915 tetanuran theropod
morphotype 3
 Theropoda - Carcharodontosauridae
Carcharodontosauridae indet.2 Stromer 1931 allosauroid
morphotype 1
 Theropoda - Elongatoolithidae
Elongatoolithidae indet. Zhao 1975 theropod
 Loricata -
Crocodyliformes indet.2 crocodilian
morphotypes 2, 4
Bernissartia sp. Dollo 1883 crocodilian
teeth
 Loricata - Krokolithidae
Krokolithes sp.1 Hirsch 1985 crocodile eggshell