Where: Orenburg, Russian Federation (51.2° N, 58.2° E: paleocoordinates 44.0° N, 51.2° E)
• coordinate stated in text
• outcrop-level geographic resolution
When: Santonian (86.3 - 83.6 Ma)
• The second smaller outcrop exposes a small section consisting, from bottom to top, of four layers, and all vertebrate material came from layer 3. Several chondrichthyan taxa that are present in the Akkermanovka assemblage were found associated with foraminifera, radiolaria, and molluscs in Santonian strata in this region, i.e., Acrolamna acuminata, Cretoxyrhina mantelli, Eostriatolamia segedini, E. venusta, Hispidaspis cf. gigas, Pseudocorax laevis, Scapanorhynchus rhaphiodon, and Squalicorax kaupi. The presence of S. kaupi in our assemblage could indicate both a Santonian or an early Campanian age for the Akkermanovka assemblage. However, none of the other taxa that are characteristic for lower Campanian strata ("Squalicorax lindstromi Zone") in the Southern Urals are present in the Akkermanovka assemblage (see Amon et al., 1997). Nonetheless, certain taxa from Akkermanovka are also reported from Campanian deposits in Russia which are outside the Southern Urals; i.e., Eostriatolamia segedini, E. venusta, Pseudocorax laevis, and Squalicorax kaupi (Averianov and Popov, 1995; Ebersole et al., 2022). Acrolamna acuminata is regarded as a typical Santonian taxon in the Southern Urals and Russia in general (Zhelezko, 1987, 1990; Amon et al., 1997), however, similar morphotypes were also described from Campanian outcrops in the Southern Urals region: i.e., A. cf. acuminata, A. acuminata dilatata (Zhelezko, 1990; Amon et al., 1997). Cretoxyrhina mantelli and Ptychodus rugosus, on the other hand, are only known from lower and upper Santonian strata of the Southern Urals (Glickman and Zhelezko, 1979, 1990; Amon et al., 1997). However, scattered reports of post-Santonian occurrences of C. mantelli exist outside Russia in Europe (Siverson, 1992b; Sørensen et al., 2013) and North America (Siverson and Lindgren, 2005; Cook et al., 2017). Based on the taxa discussed above, a Santonian-?early Campanian age is suggested for the vertebrate assemblage in this study.
Environment/lithology: coastal; poorly lithified, glauconitic, pebbly, quartzose sandstone
Size class: macrofossils
Collection methods: Repository: Museum of the World Ocean (MMO) in Kaliningrad, Russian Federation
Primary reference: P. L. Jambura, S. V. Solonin, S. L. A. Cooper, E. V. Mychko, M. S. Arkhangelsky, J. Türtscher, M. Amadori, S. Stumpf, A. V. Vodorezov and J. Kriwet. 2024. Fossil marine vertebrates (Chondrichthyes, Actinopterygii, Reptilia) from the Upper Cretaceous of Akkermanovka (Orenburg Oblast, Southern Urals, Russia). Cretaceous Research 155(105779) [M. Clapham/M. Clapham]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 232890: authorized by Matthew Clapham, entered by Matthew Clapham on 05.01.2024, edited by Matthew Carrano
Creative Commons license: CC BY (attribution)
Taxonomic list
Chondrichthyes | |
Ischyodus yanschini Averianov 1991 plownose chimaera One fragmentary left mandibular plate (MMO No 12260/ 464), one fragmentary right palatine plate (MMO No 12260/465), and one almost complete left mandibular plate (MMO No 12260/ 466)
Elasmodus sp. Egerton plownose chimaera One fragmentary right mandibular tooth plate (MMO No 12260/463)
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Chimaeroidei indet. chimaera One fragmentary left palatine plate (MMO No 12260/462) and one incomplete dorsal fin spine (MMO No 12260/467)
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Paraorthacodus cf. andersoni Case 1978 elasmobranch Twenty-four mostly complete teeth (MMO No 12260/154-177): seven anterior teeth (MMO No 12260/154, MMO No 12260/160, MMO No 12260/163, MMO No 12260/166, MMO No 12260/168, MMO No 12260/169, MMO No 12260/171), 16 lateral teeth (MMO No 12260/155-159, MMO No 12260/161-162, MMO No 12260/164-165, MMO No 12260/167, MMO No 12260/170, MMO No 12260/172-176), and one pathological tooth (MMO No 12260/177)
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Scapanorhynchus rhaphiodon Agassiz 1843 goblin shark Twenty-two teeth (MMO No 12260/198-217): six anterior teeth (MMO No 12260/199, MMO No 12260/204, MMO No 12260/ 207, MMO No 12260/211-212, MMO No 12260/217), 13 antero-lateral teeth (MMO No 12260/198, MMO No 12260/200-203, MMO No 12260/206, MMO No 12260/209-210, MMO No 12260/214-216), and three lateral teeth (MMO No 12260/205, MMO No 12260/208, MMO No 12260/213)
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Cretalamna sarcoportheta Siversson et al. 2015 mackerel shark Twenty mostly complete lateral teeth (MMO No 12260/ 75-94)
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Hispidaspis horridus Sokolov 1978 sand tiger shark Three complete anterior teeth (MMO No 12260/150, MMO No 12260/151, MMO No 12260/153)
Hispidaspis cf. gigas Woodward 1889 sand tiger shark Two complete teeth: one anterior tooth (MMO No 12260/ 149) and one antero-lateral tooth (MMO No 12260/152)
Eostriatolamia venusta Leriche 1906 sand tiger shark Thirty-two teeth (MMO No 12260/117-148): six anterior teeth (MMO No 12260/118, MMO No 12260/124, MMO No 12260/135, MMO No 12260/137, MMO No 12260/139, MMO No 12260/141), nine antero-lateral teeth (MMO No 12260/123, MMO No 12260/125-126, MMO No 12260/128, MMO No 12260/130-133, MMO No 12260/142), and 17 lateral teeth (MMO No 12260/117, MMO No 12260/119-122, MMO No 12260/127, MMO No 12260/129, MMO No 12260/134, MMO No 12260/136, MMO No 12260/138, MMO No 12260/140, MMO No 12260/143-148)
Eostriatolamia segedini Glickman and Zhelezko 1979 sand tiger shark Twenty complete teeth (MMO No 12260/97-116): four anterior teeth (MMO No 12260/97-100), five antero-lateral teeth (MMO No 12260/105-109), and eleven lateral teeth (MMO No 12260/101-104, MMO No 12260/110-116)
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Acrolamna acuminata Agassiz 1835 mackerel shark Eight mostly complete teeth (MMO No 12260/1-8): two anterior teeth (MMO No 12260/1-2), three antero-lateral teeth (MMO No 12260/3-5), and three lateral teeth (MMO No 12260/6-8)
Cretoxyrhina mantelli Agassiz 1835 mackerel shark One upper lateral tooth (MMO No 12260/95) and one lower anterior (?) tooth (MMO No 12260/96)
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Squalicorax kaupi Agassiz 1843 crow shark One hundred and nine mostly complete teeth (MMO No 12260/218-326)
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Pseudoscapanorhynchus compressidens Herman 1977 mackerel shark Thirteen well preserved teeth (MMO No 12260/185-197): ten anterior teeth (MMO No 12260/185, MMO No 12260/187, MMO No 12260/189-195, MMO No 12260/197) and three lateral teeth (MMO No 12260/186, MMO No 12260/188, MMO No 12260/197)
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Archaeolamna ex gr. kopingensis Davis 1890 mackerel shark Fifty-five teeth (MMO No 12260/9-63): ten anterior teeth (MMO No 12260/10, MMO No 12260/19, MMO No 12260/27-29, MMO No 12260/41, MMO No 12260/43, MMO No12260/49, MMO No 12260/51, MMO No 12260/53), twelve antero-lateral teeth (MMO No 12260/12, MMO No 12260/13, MMO No 12260/22-23, MMO No 12260/26, MMO No 12260/40, MMO No 12260/44, MMO No 12260/46, MMO No 12260/58, MMO No 12260/60, MMO No 12260/62-63), and 33 lateral teeth (MMO No 12260/9, MMO No 12260/11, MMO No 12260/14-18, MMO No 12260/20, MMO No 12260/21, MMO No 12260/24-25, MMO No 12260/30-39, MMO No 12260/42, MMO No 12260/45, MMO No 12260/47-48, MMO No 12260/50, MMO No 12260/51-52, MMO No 12260/55-57, MMO No 12260/59, MMO No 12260/61)
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Meristodonoides sp. Underwood and Cumbaa 2010 elasmobranch One complete and four incomplete teeth (MMO No 12260/ 333 and MMO No 12260/339-342)
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Reptilia | |
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Actinopteri | |
Lepisosteidae indet. Cuvier 1825 gar Thirteen isolated ganoid scales with varying degrees of abrasion (MMO No 12260/436-446, MMO No 12260/449, MMO No 12260/450) and two abraded dermal bone fragments (MMO No 12260/447, MMO No 12260/448)
Lepisosteidae "indet. 1" Cuvier 1825 gar Five incompletely preserved ganoid scales (MMO No 12260/436, MMO No 12260/439-441, MMO No 12260/445)
Lepisosteidae "indet. 2" Cuvier 1825 gar Four broken ganoid scales with differentiating ornamentation to morphology 1 (MMO No 12260/437, MMO No 12260/442-444)
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Protosphyraena sp. Leidy 1857 Four isolated tooth crowns of varying completeness (MMO No 12260/428-431)
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Enchodus spp. Agassiz 1835 An incomplete dermopalatine with broken tooth base (MMO No 12260/408), one partial premaxilla bone retaining dentition (MMO No 12260/435), and a broken indeterminate jaw fragment with base of broken tooth (MMO No 12260/470)
Enchodus petrosus Cope 1874 49 palatine and dermopalatine teeth in varying conditions of completeness (MMO No 12260/343-346, MMO No 12260/348-356, MMO No 12260/358-365, MMO No 12260/367-370, MMO No 12260/372-383, MMO No 12260/385, MMO No 12260/388-392, MMO No 12260/394-396, MMO No 12260/432-434)
Enchodus cf. gladiolus Cope 1872 Eight isolated tooth crowns (MMO No 12260/347, MMO No 12260/357; MMO No 12260/366, MMO No 12260/371, MMO No 12260/384, MMO No 12260/386, MMO No 12260/387, MMO No 12260/393)
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Pachyrhizodontidae indet. Cope 1872 One isolated tooth with an unusual aspect ratio compared to those reported here as cf. Pachyrhizodus sp. (MMO No 12260/412)
cf. Pachyrhizodus sp. Agassiz 1850 Eight isolated tooth crowns (MMO No 12260/411, MMO No 12260/413-419)
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