Las Hoyas Unit 3 (MCCM collection) (Cretaceous of Spain)

Where: Cuenca, Spain (40.1° N, 1.9° W: paleocoordinates 30.5° N, 8.1° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: La Huérguina Formation, Late/Upper Barremian (130.0 - 125.5 Ma)

• also called "Calizas de La Huérgina Formation" or "La Huérgina Limestones Formations"

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; limestone and packstone

• shallow to deep lacustrine facies; "produced in the context of a continental (freshwater) subtropical, seasonal summer wet, carbonate wetland that overlay a low-relief karstic terrain. The wetland was drained by carbonate-rich freshwater and comprised a typical environmental mosaic of swampy plains, ponds, lakes, ephemeral channels and sloughs."
• Limestone with laminates; three facies described: Massive limestone, packstones with peloids and sand-sized fragments of Charophyte stems. Occasionally, in the base of these beds, levels of gastropodal wackestones appear; Flaggy limestones, sometimes nodular. Ostracodal mudstones with occasional thick laminations, laterally grading to confolute laminations; Laminated limestones, mudstones with laminations containing abundant fossils. Laminae are very continuous laterally

Size classes: macrofossils, mesofossils

Preservation: adpression, trace, soft parts

Collection methods: quarrying, surface (in situ),

Primary reference: J. L. Sanz, S. Wenz, A. Yebenes, R. Estes, X. Martinez-Delclos, E. Jiminez-Fuentes, C. Dieguez, A. D. Buscalioni, L. J. Barbadillo and L. Via. 1988. An Early Cretaceous faunal and floral continental assemblage: Las Hoyas fossil site (Cuenca, Spain). Géobios 21(5):611-635 [J. Alroy/E. Leckey/J. Alroy]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 28006: authorized by John Alroy, entered by Erin Leckey on 20.01.2003, edited by Roger Benson, Matthew Carrano, John Alroy, Richard Butler, Matthew Oreska, Jered Karr, Matthew Clapham, Philip Mannion, Jonathan Tennant and Mark Bell

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Insecta
 Hemiptera -
Heteroptera indet. Latreille 1810 true bug
 Hemiptera - Naucoridae
Naucoridae indet. Fallen 1814 creeping water bug
 Hemiptera - Belostomatidae
Iberonepa romerali31 giant water bug
 Hemiptera - Notonectidae
Notonectidae indet. Latreille 1802 true bug
 Hemiptera - Corixidae
Velocorixinae indet. Popov 1986 water boatman
 Dicondylia - Chresmodidae
Chresmoda sp.31 Germar 1839 winged insect
Chresmoda aquatica6 Martínez-Delclòs 1989 winged insect
LH-26.546, LH-13826, LH-13574, LH-92AM102, MCCM-LH 18023, MCCM-LH 18024,
 Blattodea -
"Blattoidea indet." = Blattodea
"Blattoidea indet." = Blattodea Brunner von Wattenwyl 1882 cockroach
 Hymenoptera - Scoliidae
Archaeoscolia hispanica n. sp.22 Rasnitsyn and Martínez-Delclòs 1999 wasp
MCCM LH 13827
Cretoscolia conquensis n. sp.23 Rasnitsyn and Martínez-Delclòs 2000 wasp
MCCM LH 17331
 Coleoptera - Ademosynidae
Ademosynidae indet. Ponomarenko 1968 beetle
 Coleoptera - Parandrexidae
Martynopsis laticollis n. gen. n. sp.33
Martynopsis laticollis n. gen. n. sp.33 Soriano et al. 2006 beetle
MCCM LH-21091, 24513
 Coleoptera - Nemonychidae
Distenorrhinus (Parabrenthorrhinus) xavieri n. sp.38 Zherikhin and Gratshev 2003 pine flower snout beetle
MCCM LH 17330
 Coleoptera - Coptoclavidae
Hispanoclavina diazromerali n. gen. n. sp.34, Hispanoclavina gratshevi n. sp.34, Ovonectes pilosum n. gen. n. sp.34, Megacoptoclava longiurogomphia n. gen. n. sp.20, Hoyaclava buscalionae n. gen. n. sp.34
Hispanoclavina diazromerali n. gen. n. sp.34 Soriano et al. 2007 beetle
MCCM-LH16264, 15070, 16315, 22299, 23859
Hispanoclavina gratshevi n. sp.34 Soriano et al. 2007 beetle
MCCM-LH15961, 15747, 23000
Ovonectes pilosum n. gen. n. sp.34 Soriano et al. 2007 beetle
MCCM-LH23853, 24514
Megacoptoclava longiurogomphia n. gen. n. sp.20 Ponomarenko and Martínez-Delclòs 2000 beetle
MCCM LH-1263
Hoyaclava buscalionae n. gen. n. sp.34 Soriano et al. 2007 beetle
MCCM-LH24508, 7269, 23060, 24517
 Coleoptera -
Archostemata indet. Kolbe 1908 beetle
 Coleoptera - Cupedidae
Priacma sanzii n. sp.32 Soriano and Delclos 2006 reticulated beetle
MCCM LH 24524
"Anaglyphites zherikhini n. sp." = Mesocupes (Mesocupes) zherikhini32 Soriano and Delclos 2006 reticulated beetle
MCCM LH 15122
Zygadenia martinclosas32 Soriano and Delclos 2006 reticulated beetle
MCCM LH 20151
Zygadenia siniestri n. sp.32 Soriano and Delclos 2006 reticulated beetle
MCCM LH 23515, MCCM LH 21291, MCCM LH 15819
Zygadenia diazromerali n. sp.32 Soriano and Delclos 2006 reticulated beetle
MCCM LH 12001
Zygadenia longicoxa n. sp.32 Soriano and Delclos 2006 reticulated beetle
MCCM LH 928030, MCCM LH 23432
Zygadenia viridis n. sp.32 Soriano and Delclos 2006 reticulated beetle
MCCMLH 1805
Tetraphalerus ponomarenkoi n. sp.32 Soriano and Delclos 2006 reticulated beetle
MCCM LH 78
 Mecoptera - Orthophlebiidae
"Orthophlebiinae indet." = Orthophlebiidae
"Orthophlebiinae indet." = Orthophlebiidae Handlirsch 1906 scorpionfly
 Diptera - Tipulidae
Leptotarsus buscalioniae n. sp.24 Ribeiro and Lukashevich 2014 crane fly
MCCM LH 35088
Leptotarsus ibericus n. sp.24 Ribeiro and Lukashevich 2014 crane fly
MCCM LH 28206, 18122, 29995
Leptotarsus contractus n. sp.24 Ribeiro and Lukashevich 2014 crane fly
MCCM LH 15419
 Diptera - Ptychopteridae
Eoptychoptera sp.13 Handlirsch 1906 phantom crane fly
MCCM LH 35084
 Raphidioptera - Mesoraphidiidae
Mesoraphidiidae indet. Martynov 1925 snakefly
 Neuroptera - Mesochrysopidae
Allopterus mayorgai n. sp.16, "Armandochrysopa inexpecta n. sp." = Kareninoides inexpectus16, Chimerochrysopa incerta n. gen. n. sp.16, Triangulochrysopa sanzi n. gen. n. sp.16, Nanochrysopa pumilio n. gen. n. sp.16, Tachinymphes paicheleri n. sp.16, Tachinymphes penalveri n. sp.16
Allopterus mayorgai n. sp.16 Nel et al. 2005 brown lacewing
MCCM LH 18570
"Armandochrysopa inexpecta n. sp." = Kareninoides inexpectus16 Nel et al. 2005 brown lacewing
MCCM LH 18575
Chimerochrysopa incerta n. gen. n. sp.16 Nel et al. 2005 brown lacewing
MCCM LH 18588
Triangulochrysopa sanzi n. gen. n. sp.16 Nel et al. 2005 brown lacewing
MCCM LH-8110, 18572, 18573, 18574, MCCM 92/2/3
Nanochrysopa pumilio n. gen. n. sp.16 Nel et al. 2005 brown lacewing
MCCM LH 13217
Tachinymphes paicheleri n. sp.16 Nel et al. 2005 brown lacewing
MCCM LH 18576-8, LH 8040, LH 13175, MCCM LH 18579-18584
Tachinymphes penalveri n. sp.16 Nel et al. 2005 brown lacewing
MCCM LH 18585, LH 18586-7, U.A. Madrid LH 8033, 8094
 Dicondylia -
Ephemeroptera indet. Hyatt and Arms 1890 mayfly
 Odonata - Aeschnidiidae
Sinaeschnidia martinezdelclosi n. sp.11 Fleck and Nel 2003 small darner
LH 17332, ADR-317-I
Iberoaeschnidium conquensis n. gen. n. sp.11 Nel and Martínez-Delclòs 1993 small darner
LH-083-R
Gigantoaeschnidium ibericus n. gen. n. sp.11 Nel and Martínez-Delclòs 1993 small darner
Malacostraca
 Decapoda - Astacidae
Austropotamobius llopisi31 Via 1971 freshwater crayfish
 Decapoda - Oplophoridae
Bivalvia
 Unionida - Unionidae
Unio sp.31 Philipsson 1788 freshwater mussel
Gastropoda
 Cephalaspidea - Haminoeidae
Delclosia martinelli31 Rabada 1993 snail
Actinopteri
 Pholidophoriformes - Pholidophoridae
Pleuropholis sp. Egerton 1858
 Ostariophysi -
Gordichthys conquensis n. gen. n. sp.21
Gordichthys conquensis n. gen. n. sp.21 Poyato-Ariza 1994
holotype: LH 1288 R
 Gonorhynchiformes - Chanidae
 Leptolepidiformes - Leptolepididae
"? Ascalabos voithi" = Leptolepis
"? Ascalabos voithi" = Leptolepis
 Neopterygii -
Pycnodontiformes indet. Berg 1937
genus incertae sedis
 Pycnodontiformes - Pycnodontidae
 Pycnodontiformes -
Eomesodon sp.37 Woodward 1918
 Amiiformes - Caturidae
Caturus sp. Agassiz 1843 bowfin
 Amiiformes - Amiidae
Vidalamia catalunica, "? Amiopsis sp." = Amiopsis, "Urocles woodwardi" = Amiopsis
Vidalamia catalunica Sauvage 1903 bowfin
"? Amiopsis sp." = Amiopsis Kner 1863 bowfin
"Urocles woodwardi" = Amiopsis bowfin
 Lepisosteiformes -
Lepidotes sp. Agassiz 1843 gar
 Semionotiformes - Macrosemiidae
Coelacanthimorpha
 Coelacanthiformes - Latimeriidae
? Holophagus leridae coelacanth
? "Holophagus"
Osteichthyes
 Temnospondyli - Albanerpetontidae
Celtedens ibericus n. sp.15 McGowan and Evans 1995 tetrapod
LH 6020 - type (part and counterpart of a nearly complete skeleton, missing only part of the palate and tail); referred material (McGowan 2002): two complete specimens in part and counterpart (LH 030 R A & B; LH 15710 A & B)
 Salientia -
Anura indet.25, Iberobatrachus angelae n. gen. n. sp.1, Gracilibatrachus avallei n. gen. n. sp.1
Iberobatrachus angelae n. gen. n. sp.1 Baez 2013 frog
MCCM-LH 11393a-b - holotype (articulated specimen lacking the hindlimbs and left forelimb, two-dimensionally preserved as part and counterpart on two slabs); referred specimen - MCCM-LH 15999a-b (incomplete, articulated specimen preserved as part and counterpart on two slabs)
Gracilibatrachus avallei n. gen. n. sp.1 Baez 2013 frog
MCCM-LH 21171a, b - holotype (articulated skeleton of a postmetamorphic individual); referred specimens: MCCMLH-17422a, b, 11386, 11387, 26251, 26803a, b (in part), and, tentatively, 11394a, b
 Caudata -
Hylaeobatrachus sp.31 Dollo 1884 salamander
 Caudata - Urodela
Valdotriton gracilis n. gen. n. sp.10
Valdotriton gracilis n. gen. n. sp.10 Evans and Milner 1996 Wealden newt
Reptilia
 Testudines -
Hoyasemys jimenezi n. gen. n. sp.18
Hoyasemys jimenezi n. gen. n. sp.18 Pérez-García et al. 2012 turtle
MCCM-LH 84 and MCCM-LH 254 (carapace fragment)
 Theropoda -
Theropoda indet.5 theropod
tooth
 Saurischia -
? Theropoda indet.35 theropod
LH-6500, isolated track
 Avetheropoda - Carcharodontosauridae
Concavenator corcovatus n. gen. n. sp.17
Concavenator corcovatus n. gen. n. sp.17 Ortega et al. 2010 allosauroid
MCCM-LH 6666
Aves
  -
Aves indet.30 Linnaeus 1758 bird
LH 11386; 3 taxa in a pellet
 Enantiornithes -
Concornis lacustris n. gen. n. sp.28 Sanz and Buscalioni 1992 bird
LH-2814
cf. Concornis lacustris4 Sanz and Buscalioni 1992 bird
LH21006 a-b, tibiotarsus and tarsometatarsus
Eoalulavis hoyasi n. gen. n. sp.29 Sanz et al. 1996 bird
cf. Iberomesornis sp.26 Sanz et al. 1990 bird
Iberomesornis romerali n. gen. n. sp.27 Sanz et al. 1990 bird
new genus and species
Reptilia
 Avetheropoda -
Pelecanimimus polyodon n. gen. n. sp.19
Pelecanimimus polyodon n. gen. n. sp.19 Pérez-Moreno et al. 1994 ornithomimosaur
LH 7777
 Neornithischia - Iguanodontidae
Mantellisaurus atherfieldensis12 Hooley 1925 iguanodontid
LH-30677, incomplete but fully-articulated right hindlimb
 Pterosauria - Istiodactylidae
Istiodactylidae indet.35 Howse et al. 2001 pterosaur
LH-28572, LH-15448, teeth
 Pterosauria -
Europejara olcadesorum n. gen. n. sp.36
Europejara olcadesorum n. gen. n. sp.36 Vullo et al. 2012 pterosaur
MCCM-LH 9413, jaws and post-orbital elements of the skull preserved on part and counterpart slabs
 Pterosauria - Ornithocheiridae
Ornithocheiridae indet.35 Seeley 1870 pterosaur
LH-17264, LH-21451, teeth
 Loricata -
Crocodylomorpha indet.31 Hay 1930 crocodilian
 Loricata - Bernissartiidae
"Bernissartidae indet." = Bernissartiidae31
"Bernissartidae indet." = Bernissartiidae31 Dollo 1883 crocodilian
 Loricata - Atoposauridae
 Loricata -
Montsecosuchus sp.31 Buscalioni and Sanz 1988 crocodilian
 Loricata - Gobiosuchidae
Cassissuchus sanziuami n. gen. n. sp.3
Cassissuchus sanziuami n. gen. n. sp.3 Buscalioni 2017 crocodilian
 Squamata - Teiidae
Meyasaurus diazromerali n. sp.7 Evans and Barbadillo 1997 lizard
LH 370, LH 372, LH 33, LH 13510, LH 6026, LH 14317
 Squamata -
Hoyalacerta sanzi n. gen. n. sp.9, Jucaraseps grandipes n. gen. n. sp.2, Scandensia ciervensis n. gen. n. sp.8
Hoyalacerta sanzi n. gen. n. sp.9 Evans and Barbadillo 1999 lizard
LH 11000 (holotype skeleton)
Jucaraseps grandipes n. gen. n. sp.2 Bolet and Evans 2012 lizard
LH 18505 (holotype), almost complete skeleton in dorsal view
Scandensia ciervensis n. gen. n. sp.8 Evans and Barbadillo 1998 lizard
Mammalia
 Eutriconodonta - Gobiconodontidae
Spinolestes xenarthrosus n. gen. n. sp.14
Spinolestes xenarthrosus n. gen. n. sp.14 Martin et al. 2015 mammal
MCCMLH30000 - holotype (complete skeleton); MCCMLH30000B (counter slab)
Cycadeoideopsida
 Bennettitales -
Zamites sp. Brongniart 1828
Williamsonia sp. Carruthers 1870
spelled Williansonia
Pinopsida
 Pinales - Araucariaceae
Pagiophyllum sp.31 Heer 1881
 Pinales - Cheirolepidiaceae
 Pinales -
Brachyphyllum speciosa
spelled Brachyphilum
 Pinales - Taxodiaceae
Pteridopsida
  -
 Gleicheniales - Matoniaceae