Las Hoyas (MCCM collection) (Cretaceous of Spain)

Where: Castilla-La Mancha, Spain (40.1° N, 1.9° W)

• Paleocoordinates: 26.9° N, 21.3° E (Wright 2013)

• coordinate estimated from map

• outcrop-level geographic resolution

When: La Huérguina Formation, Late/Upper Barremian (125.8 - 121.4 Ma)

• also called "Calizas de La Huérgina Formation" or "La Huérgina Limestones Formations"

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; limestone and packstone

• shallow to deep lacustrine facies; "produced in the context of a continental (freshwater) subtropical, seasonal summer wet, carbonate wetland that overlay a low-relief karstic terrain. The wetland was drained by carbonate-rich freshwater and comprised a typical environmental mosaic of swampy plains, ponds, lakes, ephemeral channels and sloughs."
• Limestone with laminates; three facies described: Massive limestone, packstones with peloids and sand-sized fragments of Charophyte stems. Occasionally, in the base of these beds, levels of gastropodal wackestones appear; Flaggy limestones, sometimes nodular. Ostracodal mudstones with occasional thick laminations, laterally grading to confolute laminations; Laminated limestones, mudstones with laminations containing abundant fossils. Laminae are very continuous laterally

Size classes: macrofossils, mesofossils

Preservation: adpression, trace, soft parts

Collection methods: quarrying, surface (in situ),

• Repository: Museo de las Ciencias de Castilla-La Mancha, Cuenca (MCCM)

Primary reference: J. L. Sanz, S. Wenz, A. Yebenes, R. Estes, X. Martinez-Delclos, E. Jiminez-Fuentes, C. Dieguez, A. D. Buscalioni, L. J. Barbadillo and L. Via. 1988. An Early Cretaceous faunal and floral continental assemblage: Las Hoyas fossil site (Cuenca, Spain). Géobios 21(5):611-635 [J. Alroy/E. Leckey/J. Alroy]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 28006: authorized by John Alroy, entered by Erin Leckey on 20.01.2003, edited by Roger Benson, Matthew Carrano, John Alroy, Richard Butler, Matthew Oreska, Jered Karr, Matthew Clapham, Philip Mannion, Jonathan Tennant, Mark Bell and Bethany Allen

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names
unclassified
  -
Weitchselia reticulata, Pseudoastacus llopisi, Clavatoraxis robustus33, Turbomesodon praeclarus33, Iterophyllum lobatum33
Polypodiopsida
  -
Cladophlebis browniana
 Schizaeales - Polypodiidae
Ruffordia goepperti33
Ruffordia goepperti33 Dunker 1844
 Gleicheniales - Polypodiidae
Weichselia reticulata33
Weichselia reticulata33 Stokes and Webb 1824
Pinopsida
 Pinales - Taxodiaceae
Sphenolepis kurriana
 Bennettitales -
Williamsonia sp., Zamites sp.
Williamsonia sp. Carruthers 1870
spelled Williansonia
Zamites sp. Brongniart 1828
Angiospermae
  -
Montsechia vidali
Montsechia vidali Zeiller 1902
Equisetopsida
 Pinidae - Pinidae
Brachyphyllum speciosa, Pagiophyllum sp.33, Cupressinocladus micromerum
Brachyphyllum speciosa
spelled Brachyphilum
Pagiophyllum sp.33 Heer 1881
Actinopteri
 Neopterygii -
Pycnodontiformes indet.
Pycnodontiformes indet. Berg 1937
genus incertae sedis
 Pycnodontiformes -
Eomesodon sp.40
Eomesodon sp.40 Woodward 1918
 Pycnodontiformes - Pycnodontidae
"Coelodus subdiscus" = Ocloedus subdiscus40, Stenamara mia n. gen. n. sp.23, Macromesodon sp.40
"Coelodus subdiscus" = Ocloedus subdiscus40 Wenz 1989
Stenamara mia n. gen. n. sp.23 Poyato-Ariza and Wenz 2000
MCCM LH-14365a-b (holotype), complete skeleton
Macromesodon sp.40 Blake 1905
 Holostei - Caturidae
Caturus sp.
Caturus sp. Agassiz 1834
 Amiiformes - Amiidae
"? Amiopsis sp." = Amiopsis, "Urocles woodwardi" = Amiopsis, Vidalamia catalunica
"? Amiopsis sp." = Amiopsis Kner 1863 bowfin
"Urocles woodwardi" = Amiopsis bowfin
Vidalamia catalunica Sauvage 1903 bowfin
 Semionotiformes - Macrosemiidae
Notagogus sp., Notagogus ferreri
Notagogus sp. Agassiz 1833
 Lepisosteiformes -
Lepidotes sp., Lepidotes microrbis33, Lepidotes microrhis n. sp.39, Lepidotes tanyrhis n. sp.39
Lepidotes sp. Agassiz 1832 gar
Lepidotes microrhis n. sp.39 Wenz 2003 gar
MCCM-LH-8010a,b
Lepidotes tanyrhis n. sp.39 Wenz 2003 gar
MCCM-LH-7410
 Ostariophysi -
Gordichthys conquensis n. gen. n. sp.22
Gordichthys conquensis n. gen. n. sp.22 Poyato-Ariza 1994
holotype: LH 1288 R
 Gonorhynchiformes - Chanidae
Rubiesichthys gregalis
 Leptolepidiformes - Leptolepididae
"? Ascalabos voithi" = Leptolepis
"? Ascalabos voithi" = Leptolepis
 Pholidophoriformes - Pleuropholidae
Pleuropholis sp.
Pleuropholis sp. Egerton 1858
Reptilia
 Squamata - Teiidae
Meyasaurus diazromerali n. sp.7
Meyasaurus diazromerali n. sp.7 Evans and Barbadillo 1997 squamates
LH 370, LH 372, LH 33, LH 13510, LH 6026, LH 14317
 Squamata -
Scandensia ciervensis n. gen. n. sp.8, Jucaraseps grandipes n. gen. n. sp.2, Hoyalacerta sanzi n. gen. n. sp.9
Scandensia ciervensis n. gen. n. sp.8 Evans and Barbadillo 1998 squamates
Jucaraseps grandipes n. gen. n. sp.2 Bolet and Evans 2012 squamates
LH 18505 (holotype), almost complete skeleton in dorsal view
Hoyalacerta sanzi n. gen. n. sp.9 Evans and Barbadillo 1999 squamates
LH 11000 (holotype skeleton)
 Loricata -
Crocodylomorpha indet.33
Crocodylomorpha indet.33 Hay 1930 crocodilian
 Loricata - Atoposauridae
Atoposauridae indet.
 Loricata - Bernissartiidae
"Bernissartidae indet." = Bernissartiidae33
"Bernissartidae indet." = Bernissartiidae33 Dollo 1883 crocodilian
 Loricata -
Montsecosuchus sp.33
Montsecosuchus sp.33 Buscalioni and Sanz 1988 crocodilian
 Loricata - Gobiosuchidae
Cassissuchus sanziuami n. gen. n. sp.3
Cassissuchus sanziuami n. gen. n. sp.3 Buscalioni 2017 crocodilian
 Ornithischia -
cf. Mantellisaurus sp.15
cf. Mantellisaurus sp.15 Paul 2007 ornithopod
LH-30677, incomplete but fully-articulated right hindlimb
 Theropoda -
Theropoda indet.15, Theropoda indet.5, ? Theropoda indet.37
Theropoda indet.15 Marsh 1881 theropod
MCCM-LH-Y-1-001, trackway
Theropoda indet.5 Marsh 1881 theropod
tooth
? Theropoda indet.37 Marsh 1881 theropod
MCCM-LH-6500, isolated track
 Theropoda - Carcharodontosauridae
Concavenator corcovatus n. gen. n. sp.17
Concavenator corcovatus n. gen. n. sp.17 Ortega et al. 2010 allosauroid
MCCM-LH 6666
 Theropoda -
cf. Iberomesornis sp.28, Iberomesornis romerali n. gen. n. sp.29, Concornis lacustris n. gen. n. sp.30, cf. Concornis lacustris4, Eoalulavis hoyasi n. gen. n. sp.31
cf. Iberomesornis sp.28 Sanz et al. 1990 avialan
Iberomesornis romerali n. gen. n. sp.29 Sanz et al. 1990 avialan
new genus and species
Concornis lacustris n. gen. n. sp.30 Sanz and Buscalioni 1992 avialan
MCCM-LH-2814
cf. Concornis lacustris4 Sanz and Buscalioni 1992 avialan
MCCM-LH 21006 a-b, tibiotarsus and tarsometatarsus
Eoalulavis hoyasi n. gen. n. sp.31 Sanz et al. 1996 avialan
Aves
  -
Aves indet.32
Aves indet.32 Linnaeus 1758 bird
MCCM-LH 11386; 3 taxa in a pellet
Reptilia
 Theropoda -
Pelecanimimus polyodon n. gen. n. sp.20
Pelecanimimus polyodon n. gen. n. sp.20 Pérez-Moreno et al. 1994 ornithomimosaur
MCCM-LH 7777
 Pterosauria - Istiodactylidae
Istiodactylidae indet.37
Istiodactylidae indet.37 Howse et al. 2001 pterosaur
MCCM-LH-28572, LH-15448, teeth
 Pterosauria -
Europejara olcadesorum n. gen. n. sp.38
Europejara olcadesorum n. gen. n. sp.38 Vullo et al. 2012 pterosaur
MCCM-LH 9413, jaws and post-orbital elements of the skull preserved on part and counterpart slabs
 Pterosauria - Ornithocheiridae
Ornithocheiridae indet.37
Ornithocheiridae indet.37 pterosaur
MCCM-LH-17264, LH-21451, teeth
 Testudines -
Hoyasemys jimenezi n. gen. n. sp.19
Hoyasemys jimenezi n. gen. n. sp.19 Pérez-García et al. 2012 turtle
MCCM-LH 84, LH 254 (carapace fragment)
Mammalia
 Eutriconodonta - Gobiconodontidae
Spinolestes xenarthrosus n. gen. n. sp.13
Spinolestes xenarthrosus n. gen. n. sp.13 Martin et al. 2015 mammal
MCCM-LH 30000 - holotype (complete skeleton); MCCMLH30000B (counter slab)
Amphibia
 Temnospondyli - Albanerpetontidae
Celtedens ibericus n. sp.14
Celtedens ibericus n. sp.14 McGowan and Evans 1995 tetrapod
LH 6020 - type (part and counterpart of a nearly complete skeleton, missing only part of the palate and tail); referred material (McGowan 2002): two complete specimens in part and counterpart (LH 030 R A & B; LH 15710 A & B)
 Salientia -
Anura indet.27, Gracilibatrachus avallei n. gen. n. sp.1, Iberobatrachus angelae n. gen. n. sp.1
Anura indet.27 Fischer von Waldheim 1813 frog
Gracilibatrachus avallei n. gen. n. sp.1 Baez 2013 frog
MCCM-LH 21171a, b - holotype (articulated skeleton of a postmetamorphic individual); referred specimens: MCCMLH-17422a, b, 11386, 11387, 26251, 26803a, b (in part), and, tentatively, 11394a, b
Iberobatrachus angelae n. gen. n. sp.1 Baez 2013 frog
MCCM-LH 11393a-b - holotype (articulated specimen lacking the hindlimbs and left forelimb, two-dimensionally preserved as part and counterpart on two slabs); referred specimen - MCCM-LH 15999a-b (incomplete, articulated specimen preserved as part and counterpart on two slabs)
 Caudata -
Hylaeobatrachus sp.33
Hylaeobatrachus sp.33 Dollo 1884 salamander
 Caudata - Urodela
Valdotriton gracilis n. gen. n. sp.10
Valdotriton gracilis n. gen. n. sp.10 Evans and Milner 1996 Wealden newt
Coelacanthimorpha
 Coelacanthiformes - Latimeriidae
? Holophagus leridae
? Holophagus leridae coelacanth
? "Holophagus"
Undichna
  -
Undichna unisulca15
Undichna unisulca15 de Gibert et al. 1999
Bivalvia
 Unionida - Unionidae
Unio sp.33
Unio sp.33 Philipsson 1788 freshwater mussel
Insecta
 Mecoptera - Orthophlebiidae
"Orthophlebiinae indet." = Orthophlebiidae
"Orthophlebiinae indet." = Orthophlebiidae Handlirsch 1906 scorpionfly
 Diptera - Tipulidae
Leptotarsus contractus n. sp.26, Leptotarsus ibericus n. sp.26, Leptotarsus buscalioniae n. sp.26
Leptotarsus contractus n. sp.26 Ribeiro and Lukashevich 2014 crane fly
MCCM LH 15419
Leptotarsus ibericus n. sp.26 Ribeiro and Lukashevich 2014 crane fly
MCCM LH 28206, 18122, 29995
Leptotarsus buscalioniae n. sp.26 Ribeiro and Lukashevich 2014 crane fly
MCCM LH 35088
 Diptera - Ptychopteridae
Eoptychoptera sp.12
Eoptychoptera sp.12 Handlirsch 1906 phantom crane fly
MCCM-LH 35084
 Coleoptera - Nemonychidae
Distenorrhinus (Parabrenthorrhinus) xavieri n. sp.41
Distenorrhinus (Parabrenthorrhinus) xavieri n. sp.41 Zherikhin and Gratshev 2003 pine flower snout beetle
MCCM LH 17330
 Coleoptera - Parandrexidae
Martynopsis laticollis n. gen. n. sp.35
Martynopsis laticollis n. gen. n. sp.35 Soriano et al. 2006 beetle
MCCM LH-21091, 24513
 Coleoptera -
Archostemata indet., "Zygadenia martinclosas" = Conexicoxa martinclosas34, "Zygadenia siniestri n. sp." = Conexicoxa siniestri34, "Zygadenia diazromerali n. sp." = Notocupes diazromerali34, "Zygadenia longicoxa n. sp." = Brachilatus longicoxa34, "Zygadenia viridis n. sp." = Brachilatus viridis34
Archostemata indet. Kolbe 1908 beetle
"Zygadenia martinclosas" = Conexicoxa martinclosas34 Soriano and Delclos 2006 beetle
MCCM LH 20151
"Zygadenia siniestri n. sp." = Conexicoxa siniestri34 Soriano and Delclos 2006 beetle
MCCM LH 23515, MCCM LH 21291, MCCM LH 15819
"Zygadenia diazromerali n. sp." = Notocupes diazromerali34 Soriano and Delclos 2006 beetle
MCCM LH 12001
"Zygadenia longicoxa n. sp." = Brachilatus longicoxa34 Soriano and Delclos 2006 beetle
MCCM LH 928030, MCCM LH 23432
"Zygadenia viridis n. sp." = Brachilatus viridis34 Soriano and Delclos 2006 beetle
MCCMLH 1805
 Coleoptera - Cupedidae
Priacma sanzii n. sp.34, "Anaglyphites zherikhini n. sp." = Mesocupes zherikhini34
Priacma sanzii n. sp.34 Soriano and Delclos 2006 reticulated beetle
MCCM LH 24524
"Anaglyphites zherikhini n. sp." = Mesocupes zherikhini34 Soriano and Delclos 2006 reticulated beetle
MCCM LH 15122
 Coleoptera - Ommatidae
Tetraphalerus ponomarenkoi n. sp.34
Tetraphalerus ponomarenkoi n. sp.34 Soriano and Delclos 2006 beetle
MCCM LH 78
 Coleoptera - Ademosynidae
Ademosynidae indet.
Ademosynidae indet. Ponomarenko 1968 beetle
 Coleoptera - Coptoclavidae
Ovonectes pilosum n. gen. n. sp.36, Hoyaclava buscalionae n. gen. n. sp.36, Megacoptoclava longiurogomphia n. gen. n. sp.21, Hispanoclavina gratshevi n. sp.36, Hispanoclavina diazromerali n. gen. n. sp.36
Ovonectes pilosum n. gen. n. sp.36 Soriano et al. 2007 beetle
MCCM-LH23853, 24514
Hoyaclava buscalionae n. gen. n. sp.36 Soriano et al. 2007 beetle
MCCM-LH24508, 7269, 23060, 24517
Megacoptoclava longiurogomphia n. gen. n. sp.21 Ponomarenko and Martínez-Delclòs 2000 beetle
MCCM LH-1263
Hispanoclavina gratshevi n. sp.36 Soriano et al. 2007 beetle
MCCM-LH15961, 15747, 23000
Hispanoclavina diazromerali n. gen. n. sp.36 Soriano et al. 2007 beetle
MCCM-LH16264, 15070, 16315, 22299, 23859
 Raphidioptera - Mesoraphidiidae
Mesoraphidiidae indet.
Mesoraphidiidae indet. Martynov 1925 snakefly
 Neuroptera - Mesochrysopidae
Allopterus mayorgai n. sp.16, Triangulochrysopa sanzi n. gen. n. sp.16, "Armandochrysopa inexpecta n. sp." = Kareninoides inexpectus16, Tachinymphes paicheleri n. sp.16, Tachinymphes penalveri n. sp.16, Nanochrysopa pumilio n. gen. n. sp.16, Chimerochrysopa incerta n. gen. n. sp.16
Allopterus mayorgai n. sp.16 Nel et al. 2005 lacewing
MCCM LH 18570
Triangulochrysopa sanzi n. gen. n. sp.16 Nel et al. 2005 lacewing
MCCM LH-8110, 18572, 18573, 18574, MCCM 92/2/3
"Armandochrysopa inexpecta n. sp." = Kareninoides inexpectus16 Nel et al. 2005 lacewing
MCCM LH 18575
Tachinymphes paicheleri n. sp.16 Nel et al. 2005 lacewing
MCCM LH 18576-8, LH 8040, LH 13175, MCCM LH 18579-18584
Tachinymphes penalveri n. sp.16 Nel et al. 2005 lacewing
MCCM LH 18585, LH 18586-7, U.A. Madrid LH 8033, 8094
Nanochrysopa pumilio n. gen. n. sp.16 Nel et al. 2005 lacewing
MCCM LH 13217
Chimerochrysopa incerta n. gen. n. sp.16 Nel et al. 2005 lacewing
MCCM LH 18588
 Hymenoptera - Scoliidae
Archaeoscolia hispanica n. sp.24, Cretoscolia conquensis n. sp.25
Archaeoscolia hispanica n. sp.24 Rasnitsyn and Martínez-Delclòs 1999 hairy flower wasp
MCCM LH 13827
Cretoscolia conquensis n. sp.25 Rasnitsyn and Martínez-Delclòs 2000 hairy flower wasp
MCCM LH 17331
 Dicondylia - Chresmodidae
Chresmoda aquatica6
Chresmoda aquatica6 Martínez-Delclòs 1989 winged insect
LH-26.546, LH-13826, LH-13574, LH-92AM102, MCCM-LH 18023, MCCM-LH 18024,
 Blattodea -
"Blattoidea indet." = Blattodea
"Blattoidea indet." = Blattodea Latreille 1810 cockroach
 Hemiptera -
Heteroptera indet.
Heteroptera indet. Latreille 1810 true bug
 Hemiptera - Naucoridae
Naucoridae indet.
Naucoridae indet. Fallen 1814 creeping water bug
 Hemiptera - Belostomatidae
Iberonepa roemerali33
Iberonepa roemerali33 Martínez-Delclòs et al. 1995 giant water bug
 Hemiptera - Notonectidae
Notonectidae indet.
Notonectidae indet. Latreille 1802 true bug
 Hemiptera - Corixidae
Velocorixinae indet.
Velocorixinae indet. Popov 1986 water boatman
 Dicondylia -
Ephemeroptera indet.
Ephemeroptera indet. Hyatt and Arms 1890 mayfly
 Odonata - Aeschnidiidae
Iberoaeschnidium conquensis n. gen. n. sp.11, Gigantoaeschnidium ibericus n. gen. n. sp.11, Sinaeschnidia martinezdelclosi n. sp.11
Iberoaeschnidium conquensis n. gen. n. sp.11 Nel and Martínez-Delclòs 1993 small darner
LH-083-R
Gigantoaeschnidium ibericus n. gen. n. sp.11 Nel and Martínez-Delclòs 1993 small darner
Sinaeschnidia martinezdelclosi n. sp.11 Fleck and Nel 2003 small darner
LH 17332, ADR-317-I
Malacostraca
 Decapoda - Atyidae
Delclosia martinelli33, "Oplophorus roselli" = Delclosia roselli
Delclosia martinelli33 Rabada 1993 shrimp
"Oplophorus roselli" = Delclosia roselli Via 1971 shrimp
 Decapoda - Astacidae
Austropotamobius llopisi33
Austropotamobius llopisi33 Via 1971 freshwater crayfish
Arachnida
 Opiliones -
Opiliones indet.18
Opiliones indet.18 Sundevall 1833 harvestman
MUPA LH-29969
 Ichnofossils -
Planolites sp.15, Palaeophycus sp.15, Lockeia sp.15, Cruziana sp.15, Treptichnus sp.15, Helminthoidichnites sp.15
Planolites sp.15 Nicholson 1873
Palaeophycus sp.15 Hall 1847
Lockeia sp.15 James 1879
Cruziana sp.15 d'Orbigny 1842
Treptichnus sp.15 Miller 1889
1Baez 2013; 2Bolet and Evans 2012; 3Buscalioni 2017; 4Cambra-Moo et al. 2006; 5Canudo and Ruiz-Omeñaca 2003; 6Delclòs et al. 2008; 7Evans and Barbadillo 1997; 8Evans and Barbadillo 1998; 9Evans and Barbadillo 1999; 10Evans and Milner 1996; 11Fleck and Nel 2003; 12Lukashevich and Arillo 2016; 13Martin et al. 2015; 14McGowan and Evans 1995; 15Moratalla et al. 2017; 16Nel et al. 2005; 17Ortega et al. 2010; 18Palencia et al. 2019; 19Pérez-García et al. 2012; 20Pérez-Moreno et al. 1994; 21Ponomarenko and Martínez-Delclòs 2000; 22Poyato-Ariza 1994; 23Poyato-Ariza and Wenz 2002; 24Rasnitsyn and Martínez-Delclòs 1999; 25Rasnitsyn and Martínez-Delclòs 2000; 26Ribeiro and Lukashevich 2014; 27Sanz et al. 1988; 28Sanz and Buscalioni 1994; 29Sanz et al. 1990; 30Sanz et al. 1995; 31Sanz et al. 1996; 32Sanz et al. 2001; 33Sanz et al. 2014; 34Soriano and Delclos 2006; 35Soriano et al. 2006; 36Soriano et al. 2007; 37Vullo et al. 2009; 38Vullo et al. 2012; 39Wenz 2003; 40Wenz and Poyato-Ariza 1994; 41Zherikhin and Gratshev 2003