Sokolki locality, PIN 2005 (Permian to of Russian Federation)

Also known as Northern Dvina; Little Northern Dvina; Malaya Severnaya Dvina; Kotlas

Where: Arkhangelsk, Russian Federation (61.2° N, 46.6° E: paleocoordinates 33.5° N, 37.4° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Scutosaurus karpinskii other zone, Salarevo Formation, Late/Upper Severodvinian to Late/Upper Severodvinian (265.0 - 252.3 Ma)

• Sokolki Subassemblage of the Sokolki Assemblage; the later part of the Late Tatarian (late Severodvinian–Vyatkian) (Golubev 2000; Benton 2012, Geology).

•"Vyatka Regional Stage" (Kurkin, 1999).

•Chronostratigraphy originally entered as >>Zechstein<<.

•Lithostratigraphic data are from Ivakhnenko (2008); Salarevo Formation therein referred to as "Salarevskaya Svita" (TL).

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: macrofossils

Collected by V. P. Amalitskii; reposited in the PIN

• It appears that Amalitzky collected at the Sokolki site whereas Edemski excavated at the Zavrazhe locality (end of 19th and beginning of 20th century).

Primary reference: A. A. Kurkin. 1999. Novyy ditsinodont iz raskopok na reke Maloy Severnoy Dvine. [New dicynodont from Little Northern Dvina River excavations]. Paleontologicheskii Zhurnal 1999(3):87-92 [J. Alroy/R. Whatley/T. Liebrecht]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 28065: authorized by John Alroy, entered by Robin Whatley on 22.01.2003, edited by Torsten Liebrecht, Juan Benito Moreno and Richard Butler

Creative Commons license: CC BY (attribution)

Taxonomic list

• "Scutosaurus tuberculatus": Ivakhnenko (2008) lists PIN 2005/1578 among specimens of Scutosaurus karpinskii as well as under S. tuberculatus. Since Lee (1997) says that 2005/1578 is a juvenile and Ivakhnenko (2008) says the same about the specimen listed as S. tuberculatus is seems clear that 2005/1578 is meant to be referred to the latter taxon by Ivakhnenko (2008).

•S. karpinskii: Lee (1997) lists 9 specimens of S. karpinskii predominantly represented by single postcranial elements (PIN 2005/123, 455, 698, 1543, 1877, 2070, 2473, 2485, 2489), which are not included in the taxonomic list of the present collection.

Osteichthyes
 Tetrapoda - Chroniosuchidae
Chroniosuchus licharevi4 Riabinin 1962 tetrapod
PIN 2005/2578, 2579
Amphibia
 Temnospondyli - Dvinosauridae
"Dvinosaurus primus n. gen. n. sp." = Dvinosaurus primus8, "Dvinosaurus secundus n. sp." = Dvinosaurus primus8, "Dvinosaurus tertius n. sp." = Dvinosaurus primus8
"Dvinosaurus primus n. gen. n. sp." = Dvinosaurus primus8 Amalitzki 1921 tetrapod
PIN 2005/39 (holotype), skull and anterior postcranial skeleton
"Dvinosaurus secundus n. sp." = Dvinosaurus primus8 Amalitzki 1921 tetrapod
PIN 2005/40 (holotype)
"Dvinosaurus tertius n. sp." = Dvinosaurus primus8 Amalitzki 1921 tetrapod
PIN 2005/41 (holotype)
Osteichthyes
 Therapsida - Inostranceviidae
"Inostrancevia alexandri n. gen. n. sp." = Inostrancevia alexandri5, "Inostrancevia proclivis n. sp." = Inostrancevia alexandri5, "Inostrancevia latifrons n. sp." = Inostrancevia latifrons5, "Amalitzkia vladimiri n. gen. n. sp." = Inostrancevia latifrons5
"Inostrancevia alexandri n. gen. n. sp." = Inostrancevia alexandri5 Amalitski 1922 therapsid
PIN 2005/1587 (lectotype), skeleton. PIN 2005/1588, skeleton, PIN 2005/2099, skull roof
"Inostrancevia proclivis n. sp." = Inostrancevia alexandri5 Amalitski 1922 therapsid
PIN 2005/1858 (type)
"Inostrancevia latifrons n. sp." = Inostrancevia latifrons5 Pravoslavlev 1927 therapsid
PIN 2005/1857 (type)
"Amalitzkia vladimiri n. gen. n. sp." = Inostrancevia latifrons5 Pravoslavlev 1927 therapsid
PIN 2005/1856 (type)
 Therapsida - Gorgonopidae
"Inostrancevia parva n. sp." = Pravoslavlevia parva5
"Inostrancevia parva n. sp." = Pravoslavlevia parva5 Pravoslavlev 1927 therapsid
PIN 2005/1859 (holotype), skull
 Dicynodontia -
"Dicynodon amalitzkii n. sp." = Peramodon amalitzkii9, "Dicynodon trautscholdi n. sp." = Vivaxosaurus trautscholdi1, "Gordonia annae n. sp." = Vivaxosaurus trautscholdi1, "Gordonia rossica n. sp." = Vivaxosaurus trautscholdi1, "Oudenodon venyukovi n. sp." = Vivaxosaurus trautscholdi1
"Dicynodon amalitzkii n. sp." = Peramodon amalitzkii9 Seeley 1908 dicynodont
PIN 2005/38, a well preserved complete skull and lower jaws, axial column (missing the caudal region), pectoral girdle and fragmentary pelvic girdle
"Dicynodon trautscholdi n. sp." = Vivaxosaurus trautscholdi1 Amalitzky 1922 dicynodont
PIN 2005/1 a well preserved, complete skull and lower jaws
"Gordonia annae n. sp." = Vivaxosaurus trautscholdi1 Amalitzky 1922 dicynodont
PIN 2005/5 a poorly preserved but nearly complete skull missing portions of the squamosals
"Gordonia rossica n. sp." = Vivaxosaurus trautscholdi1 Amalitzky 1922 dicynodont
PIN 2005/3 a badly distorted skull
"Oudenodon venyukovi n. sp." = Vivaxosaurus trautscholdi1 Amalitzky 1922 dicynodont
PIN 2005/6 a poorly preserved, laterally crushed skull
 Dicynodontia - Dicynodontidae
Elph borealis Kurkin 1999 dicynodont
PIN 2005/8, 2005/2643
 Therapsida - Dviniidae
"Dvinia prima n. gen. n. sp." = Dvinia prima2, "Permocynodon sushkini n. gen. n. sp." = Dvinia prima2
"Dvinia prima n. gen. n. sp." = Dvinia prima2 Amalitzky 1922 cynodont
PIN 2005/2465 (holotype), anterior half of skull. PIN 2245/1237, a right maxilla
"Permocynodon sushkini n. gen. n. sp." = Dvinia prima2 Amalitzky 1922 cynodont
PIN 2005/2469 (holotype), almost complete skull
 Therocephalia - Annatherapsididae
"Anna petri n. gen. n. sp." = Annatherapsidus petri7
"Anna petri n. gen. n. sp." = Annatherapsidus petri7 Amalitzky 1922 therapsid
PIN 2005/1993 (holotype), partial skull. PIN 2005/1562, 1991, 1992, 1994, 1995, 1996, 1997, 1998, 2155
Reptilia
 Procolophonomorpha - Pareiasauridae
"Pareiosaurus karpinskii n. sp." = Scutosaurus karpinskii6, "Scutosaurus karpinskii" = Scutosaurus karpinskii6, "Pareiosaurus tuberculatus n. sp." = Scutosaurus karpinskii6, "Scutosaurus tuberculatus" = Scutosaurus karpinskii6, "Pareiosaurus elegans n. sp." = Scutosaurus karpinskii6, "Pareiosaurus horridus n. sp." = Scutosaurus karpinskii6
"Pareiosaurus karpinskii n. sp." = Scutosaurus karpinskii6 Amalitzky 1922 parareptile
PIN 2005/1532 (type)
"Scutosaurus karpinskii" = Scutosaurus karpinskii6 Amalitzky 1922 parareptile
PIN 2005/1534 (a skull, listed by Lee, 1997, ?erroneously listed as 2005/1543 in Ivakhnenko, 2008), 2005/1536*, 37*, 2005/1573**, 2005/1883**, 2005/1885, 2005/2471, 72**, 2005/2546**; all are represented by skeletons or skulls; *mark specimens mentioned in Lee, 1997, but not in Ivakhnenko, 2008; **mark specimens not mentioned in Lee, 1997, but in Ivakhnenko, 2008; for additonal material listed by Lee, 1997, see comments on taxonomic list
"Pareiosaurus tuberculatus n. sp." = Scutosaurus karpinskii6 Amalitzky 1922 parareptile
PIN 2005/1533 (type)
"Scutosaurus tuberculatus" = Scutosaurus karpinskii6 Amalitzky 1922 parareptile
PIN 2005/1578 (a juvenile); see comments on taxonomic list
"Pareiosaurus elegans n. sp." = Scutosaurus karpinskii6 Amalitzky 1922 parareptile
PIN 2005/1538 (type)
"Pareiosaurus horridus n. sp." = Scutosaurus karpinskii6 Amalitzky 1922 parareptile
PIN 2005/1535 (type)
Osteichthyes
 Seymouriamorpha - Karpinskiosauridae
"Kotlassia secunda n. sp." = Karpinskiosaurus secundus3
"Kotlassia secunda n. sp." = Karpinskiosaurus secundus3 Amaltzki 1898 tetrapod
PIN 2005/81 (holotype), 82
 Seymouriamorpha - Kotlassidae
Kotlassia prima n. gen. n. sp.3
Kotlassia prima n. gen. n. sp.3 Amalitzky 1898 tetrapod
PIN 2005/74 (holotype). PIN 2005/75, 2689