Vero (Pleistocene of the United States)

Where: Indian River County, Florida (27.6° N, 80.4° W: paleocoordinates 27.6° N, 80.4° W)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Late/Upper Pleistocene (0.1 - 0.0 Ma)

• Stratum 3 of Vero excavation produced most materials (= "muck bed"); Stratum 2 less productive but supposedly not significantly older

•plants are mostly from layer No. 3 and vertebrates are mostly from layer No. 2 according to Hrdlicka 1917

• bed-level stratigraphic resolution

Environment/lithology: "channel"; poorly lithified sandstone and unlithified, white, carbonaceous sandstone

• "The lower sand [= Stratum 2] marks the recession of the sea in which the underlying shell marl was formed. The upper beds (No. 3) mark successive seasonal layers of valley filling in the narrow valley of a small stream. The stream was apparently always small."
• "The former [human skeleton] lay in dark and somewhat indurated sands, layer No. 2 of Sellards, the latter for the most part at the base of layer No. 3, the muck deposit of the stream bed... layer No. 2 is seen to be overlain by a stratum of similar, but somewhat lighter, sandy deposits covered by a layer of marl [ranging] from about 5 to 9 inches in thickness, and when freshly exposed is of the consistency of fresh mortar, but on exposure hardens to fairly solid rock... The second skeleton lay... in some loose white sand and vegetable matter at the base of the muck layer" (Hrdlicka 1917)

Size classes: macrofossils, mesofossils

Collected in 1913

Collection methods: quarrying, mechanical,

Primary reference: R. W. Shufeldt. 1917. Fossil birds found at Vero, Florida. Florida State Geological Survey Annual Report 9:35-42 [M. Carrano/M. Carrano/M. Uhen]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 34269: authorized by Matthew Carrano, entered by Matthew Carrano on 25.08.2003, edited by Richard Hulbert, Evangelos Vlachos, John Alroy, Austin Hendy and Uta Merkel

Creative Commons license: CC BY (attribution)

Taxonomic list

Pinopsida
 Pinales - Pinaceae
Pinus sp.2 Linnaeus 1753 pine
Pinus taeda2 Linnaeus 1753 pine
Pinus caribaea2 Morelet 1855 pine
 Coniferales - Cupressaceae
Taxodium distichum2 Linnaeus 1753 bald cypress
Angiospermae
 Laurales - Lauraceae
 Poales - Cyperaceae
Carex sp.2 Linnaeus 1753 sedge
 Mesangiosperms - Araceae
Pistia spathulata2 Michaux 1803 water cabbage
 Mesangiosperms - Arecaceae
Serenoa serrulata2 Michaux 1803 fan palm
Sabal palmetto2 Walter 1788 palm
 Asterales - Asteraceae
Xanthium sp.2 Linnaeus 1753 cocklebur
 Aquifoliales - Aquifoliaceae
Ilex glabra2 Gray 1856 holly
 Dipsacales - Caprifoliaceae
Viburnum cf. dentatum2 Linnaeus 1753 honeysuckle
Viburnum nudum2 Linnaeus 1753 honeysuckle
 Caryophyllales - Polygonaceae
Polygonum sp.2 Linnaeus 1753 knotweed
 Nymphaeales - Nymphaeaceae
Brasenia purpurea2 Caspary 1873 water lily
 Magnoliales - Annonaceae
Anona glabra2 Linnaeus 1753 custard apple
 Magnoliales - Magnoliaceae
Magnolia virginiana2 Linnaeus 1753 magnolia
 Fagales - Myricaceae
Myrica cerifera2 Linnaeus 1753 bayberry
 Fagales - Fagaceae
Quercus brevifolia2 Sargent 1895 oak
Quercus ? chapmani2 Sargent 1895 oak
Quercus virginiana2 Miller 1768 oak
Quercus laurifolia2 Michaux 1801 oak
 Rosales - Rhamnaceae
Zizyphus sp.2 Miller 1758 buckthorn
 Sapindales - Sapindaceae
Acer rubrum2 Linnaeus 1753 maple
 Vitales - Vitaceae
Vitis sp.2 Linnaeus 1753 grapevine
Vitis cf. rotundifolia2 Michaux 1803 grapevine
Leitneria floridana
  -
Leitneria floridana2 Chapman 1860 corkwood
Anthozoa
 Scleractinia - Oculinidae
Oculina robusta11 Pourtales 1871 stony coral
 Scleractinia - Faviidae
Solenastrea hyades8 Dana 1846 stony coral
Chondrichthyes
 Myliobatiformes - Aetobatidae
"Aetobatis narinari" = Aetobatus5
"Aetobatis narinari" = Aetobatus5 stingray
Actinopteri
 Lepisosteiformes - Lepisosteidae
 Amiiformes - Amiidae
"Amiatus calvus" = Amia5
"Amiatus calvus" = Amia5 bowfin
= Amia calva
 Carangiformes - Carangidae
Caranx sp.5 Lacépède 1802
not C. hippos
Reptilia
 Testudines - Cheloniidae
Caretta caretta5 Linnaeus 1758 loggerhead sea turtle
Chelonia mydas5 Linnaeus 1758 green turtle
 Testudines - Testudinidae
Gopherus polyphemus5, "Testudo sellardsi n. sp." = Hesperotestudo crassiscutata4, Testudo luciae n. sp.4, Gopherus praecedens n. sp.4
Gopherus polyphemus5 Daudin 1802 Florida gopher tortoise
"Testudo sellardsi n. sp." = Hesperotestudo crassiscutata4 Leidy 1889 turtle
USNM 8817 (holotype), a left xiphiplastron (Hay 1916a, pl. 8.6–8; Auffenberg 1963, fig. 17).
Testudo luciae n. sp.4 Hay 1916 turtle
USNM 8818 (holotype), a partial right hypoplastron (Hay 1916a, pl. 9.5; Auffenberg 1963, fig. 17).
Gopherus praecedens n. sp.4 Hay 1916 turtle
USNM 8828 (holotype), a left xiphiplastron (Hay 1916a, pl. 4.1–2), formerly cataloged under FGS 5463
 Testudines - Emydidae
Pseudemys floridana5, Trachemys sculpta4, "Trachemys bisornata" = Pseudemys bisornatus4, "Terrapene innoxia" = Terrapene carolina5, "Terrapene canaliculata" = Terrapene putnami3, "Terrapene antipex" = Terrapene putnami5
Pseudemys floridana5 Le Conte 1830 peninsula cooter
"Pseudemys floridianus persimilis"
Trachemys sculpta4 Hay 1908 slider
"Trachemys bisornata" = Pseudemys bisornatus4 Cope 1877 turtle
distal end of a fifth costal
"Terrapene innoxia" = Terrapene carolina5 Linnaeus 1758 box turtle
many specimens including complete carapaces
"Terrapene canaliculata" = Terrapene putnami3 Hay 1906 box turtle
USNM 11330, AM 25-144
"Terrapene antipex" = Terrapene putnami5 Hay 1906 box turtle
many specimens; single bones
 Testudines -
? Trachemys nuchocarinata5 Hay 1916 turtle
 Testudines - Chelydridae
"Chelydra laticarinata n. sp." = Chelydra serpentina4, "Chelydra sculpta" = Chelydra serpentina5, "Chelydra sculpta n. sp." = Chelydra serpentina4
"Chelydra laticarinata n. sp." = Chelydra serpentina4 Linnaeus 1758 Common snapping turtle
USNM V8827 (holotype, formerly FGS 7094), an isolated left peripheral VI (Hay 1916, pl. 6.6, 7)
"Chelydra sculpta" = Chelydra serpentina5 Linnaeus 1758 Common snapping turtle
"Chelydra sculpta n. sp." = Chelydra serpentina4 Linnaeus 1758 Common snapping turtle
USNM V8826 (holotype, formerly FGS 5510), an isolated right peripheral IX
Aves
  -
Aves indet. Linnaeus 1758 bird
Stratum 3
 Charadriiformes - Laridae
? Larus ? sp. Linnaeus 1758 gull
Stratum 3
 Anseriformes - Anatidae
Querquedula floridiana n. sp. Shufeldt 1917 duck
Stratum 2
 Strigiformes - Tytonidae
"Tyto pratincola" = Tyto alba pratincola
"Tyto pratincola" = Tyto alba pratincola Bonaparte 1838 barn owl
Stratum 3
 Cathartiformes - Cathartidae
Cathartes aura Linnaeus 1758 turkey vulture
Stratum 2
 Ciconiiformes - Ciconiidae
"Jabiru weillsi n. sp." = Jabiru mycteria10, Mycteria ? americana
"Jabiru weillsi n. sp." = Jabiru mycteria10 Lichtenstein 1819 jabiru
Mycteria ? americana Linnaeus 1758 wood stork
Stratum 3
 Pelecaniformes - Ardeidae
? Ardeidae ? indet. Leach 1820 heron
"comes nearest to Herodias egretta"
? Ardea ? sp. Linnaeus 1758 heron
Ardea? (sp.?); Stratum 3
"Herodias egretta" = Ardea alba egretta Gmelin 1789 great egret
Ardea herodias Linnaeus 1758 great blue heron
Stratum 3
"Larus vero n. sp." = Nyctanassa violacea Linnaeus 1758 yellow-crowned night heron
Stratum 3
 Suliformes - Phalacrocoracidae
Phalacrocorax auritus13 Lesson 1831 double-crested cormorant
Reptilia
 Galliformes - Phasianidae
"Meleagris gallopavo" = Meleagris gallopavo13, "Ardea sellardsi n. sp." = Meleagris gallopavo
"Meleagris gallopavo" = Meleagris gallopavo13 Linnaeus 1758 wild turkey
"Ardea sellardsi n. sp." = Meleagris gallopavo Linnaeus 1758 wild turkey
Stratum 3
 Crocodylia - Alligatoridae
Alligator mississippiensis5 Daudin 1801 American alligator
 Squamata - Crotalidae
Crotalus adamanteus5 Palisot de Beauvois 1799 eastern diamondback rattlesnake
Gilmore 1938 (14699): USNM 11333, 46 thoracic vertebrae; Fla. Geol Surv. No. V 2411 and 2412, each 2 vertebrae; V 2414, 2399, 1702, 191, 1712, 1765 single vertebrae each
 Squamata - Colubridae
Drymarchon corais5 Boie 1827 cribo
Gilmore 1938 (14699): (Drymarchon cf. corais couperi) Florida Geol. Surv. No. V 1630, two thoracic vertebrae; V 1546, three thoracic vertebrae; V 2163, two vertebrae
Farancia abacura5 Holbrook 1836 colubrid snake
MATERIAL LOST. Gilmore 1938 (14699): "The failure to find among the Florida serpent materials any vertebrae which can be referred to the genus Farancia leads me to question the authenticity of Hay’s reference of a single articular bone, no longer available, to F. abacura."
Mammalia
 Didelphimorphia - Didelphidae
Didelphis virginiana5 Kerr 1792 Virginia opossum
 Proboscidea - Mammutidae
Mammut americanum5 Kerr 1792 American mastodon
Stratum 3
 Proboscidea - Elephantidae
Mammuthus columbi1 Falconer 1857 Columbian mammoth
 Primates - Hominidae
Homo sapiens11 Linnaeus 1758 human
 Rodentia - Cricetidae
Sigmodon hispidus5 Say and Ord 1825 hispid cotton rat
Neotoma floridana5 Ord 1818 eastern woodrat
Reithrodontomys humulis12 Audubon and Bachman 1841 eastern harvest mouse
Oryzomys ? palustris5 Harlan 1837 marsh rice rat
Neofiber alleni5 True 1884 round-tailed muskrat
 Lagomorpha - Leporidae
Sylvilagus palustris5 Bachman 1837 marsh rabbit
Stratum 3
 Soricomorpha - Talpidae
Scalopus aquaticus5 Linnaeus 1758 eastern mole
subsp. = australis
 Artiodactyla - Tayassuidae
Mylohyus fossilis1 Leidy 1860 peccary
Strata 2 and 3
 Artiodactyla - Cervidae
Odocoileus sellardsiae n. sp.5 Hay 1917 New World deer
Odocoileus virginianus9 Zimmermann 1780 white-tailed deer
 Artiodactyla - Bovidae
Bison sp.5 Hamilton-Smith 1827 bison
Stratum 3
 Perissodactyla - Equidae
Equus sp.1 Linnaeus 1758 horse
 Perissodactyla - Tapiridae
"Tapirus veroensis n. sp." = Tapirus (Helicotapirus) veroensis7
"Tapirus veroensis n. sp." = Tapirus (Helicotapirus) veroensis7 Sellards 1918 tapir
 Carnivora - Canidae
"Vulpes palmaria n. sp." = Vulpes vulpes5, Canis sp.5, Canis latrans1, "Canis ayersi n. sp." = Aenocyon dirus10
"Vulpes palmaria n. sp." = Vulpes vulpes5 Linnaeus 1758 red fox
Canis sp.5 Linnaeus 1758 canine
Stratum 3
Canis latrans1 Say 1823 coyote
"Canis ayersi n. sp." = Aenocyon dirus10 Leidy 1858 dire wolf
 Carnivora - Felidae
Lynx rufus5 Schreber 1777 bobcat
subsp. = floridianus
 Carnivora - Mustelidae
Lontra canadensis1 Schreber 1776 North American river otter
 Carnivora - Procyonidae
Procyon lotor5 Linnaeus 1758 raccoon
 Carnivora - Ursidae
Ursus americanus1 Pallas 1780 black bear
 Cingulata - Pampatheriidae
Holmesina septentrionalis1 Leidy 1889 edentate
 Cingulata - Dasypodidae
Dasypus sp.5 Linnaeus 1758 armadillo
Stratum 3
Amphibia
 Caudata - Urodela
Amphiuma means5 Garden 1821 two-toed amphiuma
Siren lacertina5 Linnaeus 1766 greater siren
Gymnolaemata
 Cheilostomata - Microporidae
Cupularia denticulata11 Conrad 1841
Gastropoda
 Lepetellida - Fissurellidae
"Fissurella alternata" = Diodora cayenensis11
"Fissurella alternata" = Diodora cayenensis11 Lamarck 1822 snail
 Cerithioidea - Turritellidae
Vermicularia nigricana11 turret shell
Turritella sp.11 Lamarck 1799 turret shell
 Cerithioidea - Cerithiidae
Cerithium sp.11, Cerithium aff. muscarum11, "Cerithium floridanum" = Cerithium atratum11
Cerithium sp.11 Bruguière 1789 cerith snail
Cerithium aff. muscarum11 Say 1832 cerith snail
"Cerithium floridanum" = Cerithium atratum11 Born 1778 cerith snail
 Cerithioidea - Modulidae
"Modulus floridanus" = Modulus modulus11
"Modulus floridanus" = Modulus modulus11 Linnaeus 1758 snail
 Sorbeoconcha - Naticidae
"Sigaretus perspectivus" = Sinum perspectivum11, "Neverita duplicata" = Neverita duplicatus11
"Sigaretus perspectivus" = Sinum perspectivum11 Say 1831 moon snail
"Neverita duplicata" = Neverita duplicatus11 Say 1822 moon snail
 Sorbeoconcha - Littorinidae
"Littorina irrorata" = Littoraria irrorata11
"Littorina irrorata" = Littoraria irrorata11 Say 1822 periwinkle
 Sorbeoconcha - Vermetidae
"Petaloconchus irregularis" = Dendropoma irregulare11
"Petaloconchus irregularis" = Dendropoma irregulare11 d'Orbigny 1841 worm shell
 Neogastropoda - Columbellidae
"Columbella avara" = Costoanachis avara11
"Columbella avara" = Costoanachis avara11 Say 1822 snail
 Neogastropoda - Fasciolariidae
"Fasciolaria distans" = Cinctura lilium11
"Fasciolaria distans" = Cinctura lilium11 Fischer 1807 snail
 Neogastropoda - Nassariidae
"Nassa acuta" = Nassarius (Nassarius) acutus11
"Nassa acuta" = Nassarius (Nassarius) acutus11 Say 1822 snail
 Neogastropoda - Melongenidae
Busycon perversum11 Linnaeus 1758 whelk
Busycon carica11 Gmelin 1792 whelk
Melongena corona11 Gmelin 1791 snail
 Neogastropoda - Marginellidae
"Marginella aff. apicina" = Prunum apicinum11, Marginella sp.11
"Marginella aff. apicina" = Prunum apicinum11 Menke 1828 margin shell
Marginella sp.11 Lamarck 1799 margin shell
 Neogastropoda - Muricidae
Murex fulvenscens11 murex snail
Purpura haemostoma11 murex snail
Eupleura caudata11 Say 1822 murex snail
? Urosalpinx sp.11 Stimpson 1865 murex snail
 Neogastropoda - Olividae
"Olivella ? jaspidea" = Jaspidella jaspidea11, "Oliva literata" = Oliva (Ispidula) sayana11, Olivella mutica11
"Olivella ? jaspidea" = Jaspidella jaspidea11 Gmelin 1791 olive snail
"Oliva literata" = Oliva (Ispidula) sayana11 Ravenel 1834 olive snail
Olivella mutica11 Say 1822 olive snail
 Neogastropoda - Terebridae
"Terebra dislocata" = Strioterebrum dislocatum11, "Terebra concava" = Strioterebrum concava11, "Terebra cinerea" = Impages cinerea11
"Terebra dislocata" = Strioterebrum dislocatum11 Say 1822 auger snail
"Terebra concava" = Strioterebrum concava11 Say 1827 auger snail
"Terebra cinerea" = Impages cinerea11 Born 1778 auger snail
 Neotaenioglossa - Calyptraeidae
Crepidula plana11 Say 1822 slipper shell
Crepidula fornicata11 Linnaeus 1758 slipper shell
 Epitonioidea - Epitoniidae
Epitonium lineata11 wentletrap
Epitonium sp.11 Bolten 1798 wentletrap
Bivalvia
 Nuculida - Nuculidae
"Nucula proxima" = Nucula (Nucula) proxima11
"Nucula proxima" = Nucula (Nucula) proxima11 Say 1820 Atlantic Nut Clam
 Arcida - Arcidae
Arca (Noetia) limula11, Acar domingensis6, "Arca cf. umbonata" = Lamarcka imbricata11, "Scapharca (Scapharca) transversa" = Anadara (Larkinia) transversa11, "Scapharca (Argina) campechiensis" = Anadara (Lunarca)11, "Scapharca (Cunearca) incongrus" = Anadara (Cunearca)11
Arca (Noetia) limula11 Conrad 1832 ark
Acar domingensis6 Lamarck 1819 miniature ark
"Arca cf. umbonata" = Lamarcka imbricata11 Bruguière 1789 clam
"Scapharca (Scapharca) transversa" = Anadara (Larkinia) transversa11 Say 1822 ark
"Scapharca (Argina) campechiensis" = Anadara (Lunarca)11 ark
"Scapharca (Cunearca) incongrus" = Anadara (Cunearca)11 ark
 Arcida - Glycymerididae
Glycymeris americana11, "Glycymeris pectinata" = Tucetona pectinata11
Glycymeris americana11 Defrance 1826 clam
"Glycymeris pectinata" = Tucetona pectinata11 Gmelin 1791 clam
 Pectinida - Anomiidae
Anomia simplex11 d'Orbigny 1845 jingle
 Pectinida - Plicatulidae
 Pectinida - Pectinoidae
"Pecten gibbus" = Argopecten gibbus11
"Pecten gibbus" = Argopecten gibbus11 Linnaeus 1758 calico scallop
 Carditida - Crassatellidae
Crassinella sp.11 Guppy 1874 clam
Crassinella ? lunulata11 Conrad 1834 clam
 Carditida - Carditidae
"Venericardia tridentata" = Pleuromeris tridentata11, "Venericardia perplana" = Pteromeris perplana11, "Cardita (Carditamera) floridana" = Carditamera floridana11
"Venericardia tridentata" = Pleuromeris tridentata11 Say 1826 clam
"Venericardia perplana" = Pteromeris perplana11 Dall 1903 clam
"Cardita (Carditamera) floridana" = Carditamera floridana11 Conrad 1838 clam
 Lucinida - Lucinidae
"Phacoides (Parvilucina) multilineatus" = Parvilucina multistriata11, Phacoides aff. nasuta11, Phacoides sp.11, "Phacoides floridanus" = Stewartia floridana11, "Divaricella quadrisulcata" = Divalinga quadrisulcata11
"Phacoides (Parvilucina) multilineatus" = Parvilucina multistriata11 Conrad 1841 clam
Phacoides sp.11 Blainville 1825 clam
"Phacoides floridanus" = Stewartia floridana11 Conrad 1833 clam
"Divaricella quadrisulcata" = Divalinga quadrisulcata11 d'Orbigny 1846 clam
 Pholadida - Corbulidae
"Corbula ? cuneata" = Caryocorbula cuneata11
"Corbula ? cuneata" = Caryocorbula cuneata11 Say 1824 clam
 Cardiida - Donacidae
Donax tumida11, "Donax variabilis" = Serrula variabilis11
"Donax variabilis" = Serrula variabilis11 Say 1822 clam
 Cardiida - Semelidae
Semele proficua11 Pulteney 1799 clam
Semele ? bellastriata11 Conrad 1837 clam
 Cardiida - Tellinidae
Strigilla sp.11, "Strigilla flexuosa" = Strigilla mirabilis11
Strigilla sp.11 Turton 1822 tellin clam
"Strigilla flexuosa" = Strigilla mirabilis11 Philippi 1841 tellin clam
 Cardiida - Cardiidae
Cardium (Laevicardium) mortoni11, "Cardium (Laevicardium) serratum" = Laevicardium serratum11, "Cardium (Trachycardium) isocardia" = Trachycardium (Trachycardium) isocardia11, "Cardium (Trachycardium) muricatum" = Trachycardium (Dallocardia) muricatum11, "Cardium (Cerastoderma) robustum" = Dinocardium robustum11
Cardium (Laevicardium) mortoni11 Conrad 1831 cockle
"Cardium (Laevicardium) serratum" = Laevicardium serratum11 Linnaeus 1758 cockle
"Cardium (Trachycardium) isocardia" = Trachycardium (Trachycardium) isocardia11 Linnaeus 1758 cockle
"Cardium (Trachycardium) muricatum" = Trachycardium (Dallocardia) muricatum11 Linnaeus 1758 cockle
"Cardium (Cerastoderma) robustum" = Dinocardium robustum11 Lightfoot 1786 cockle
 Cardiida - Mactridae
Mactra sp.11 Linnaeus 1767 clam
"Spisula solidissima" = Hemimactra solidissima11 Dilwyn 1817 clam
Mulinia lateralis11 Say 1822 clam
 Cardiida - Veneridae
Venus sp.11, "Venus campechiensis" = Mercenaria campechiensis11, "Venus mortoni" = Mercenaria campechiensis11, Chione interpurpurea11, Chione sp.11, Chione cancellata11, "Chione (Lirophora) latilirata" = Lirophora (Lirophora) latilirata11, Anomalocardia cuneimeris11, "Chione (Timoclea) grus" = Timoclea grus11, Dosinia distans11, Dosinia sp.11, Transennella caloosana11
Venus sp.11 Linnaeus 1758 venus clam
"Venus campechiensis" = Mercenaria campechiensis11 Gmelin 1791 quahog
"Venus mortoni" = Mercenaria campechiensis11 Gmelin 1791 quahog
Chione interpurpurea11 venus clam
Chione sp.11 von Mühlfield 1811 venus clam
Chione cancellata11 Linnaeus 1767 venus clam
"Chione (Lirophora) latilirata" = Lirophora (Lirophora) latilirata11 Conrad 1841 venus clam
Anomalocardia cuneimeris11 Conrad 1846 venus clam
"Chione (Timoclea) grus" = Timoclea grus11 Holmes 1858 venus clam
Dosinia distans11 venus clam
Dosinia sp.11 Scopoli 1777 venus clam
Merisca aequistriata
  -
"Tellina lintea" = Merisca aequistriata11
"Tellina lintea" = Merisca aequistriata11 Say 1824