Eggingen-Mittelhart 1 (Oligocene of Germany)

Where: Baden-Württemberg, Germany (48.4° N, 9.9° E: paleocoordinates 48.2° N, 7.0° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Ulmer Schichten Member (Lower Freshwater Molasse Formation), MP 30 (28.4 - 23.0 Ma)

• basal "Ulm beds", Oligocene/Miocene transition (Werner, 1994 p. 9)

•Werner (1994, p. 202), however, says that only Eggingen-Mittelhart 2 provided a mammal fauna that is transitional between the Upper Oligocene and the Lower Miocene whereas Eggingen-Mittelhart 1 can clearly be dated as Upper Oligocene (MP 30).

•stratigraphy, originally entered as >>Late/Upper Oligocene<< to >>Early/Lower Miocene<< has been modified accordingly (TL)

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine; poorly lithified, shelly/skeletal, black, brown, calcareous, carbonaceous mudstone and unlithified sandstone

• The locality is in the so called "Ulmer Schichten" ("Ulm beds"), wich represent the Lower Freshwater Molasse in the Ulm area, the latter being part of the northern margin of the alpine molasse basin. The "Ulm beds" are also referred to as "subjurassic molasse" due to its position immediately south of the Swabian Jura (Schwäbische Alb) (Werner, 1994).
• "Graupensande" of the Brackwater Molasse (MN4b) cut down into deep brown to black, marly, highly fossiliferous bituminous mudstones ('chocolate marls') of the Lower Freshwater Molasse, the latter being partly interspersed with gastropod shell debris (Werner, 1994)

Size class: mesofossils

Preservation: original phosphate

Collected in 1983; reposited in the BSP, SMNS

Collection methods: bulk, sieve,

• The fossiliferous layer is not accessible anymore (Werner, 1994).

•EGMH-1 refers to material housed in the Wannemacher collection

•Original total collection size entry (4500 specimens) has been removed since individual specimen numbers have been entered for each taxon. (TL)

Primary reference: J. Werner. 1994. Beiträge zur Biostratigraphie der Unteren Süßwasser-Molasse Süddeutschlands - Rodentia und Lagomorpha (Mammalia) aus den Fundstellen der Ulmer Gegend. Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie). 200:1-263 [L. van den Hoek Ostende/L. van den Hoek Ostende/L. van den Hoek Ostende]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 66109: authorized by Lars van den Hoek Ostende, entered by Lars van den Hoek Ostende on 28.09.2006, edited by Juliette Richter, Torsten Liebrecht and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

• The individual numbers of teeth of Rhodanomys transiens and R. latens are not given in the reference. Instead it is said, that 2836 teeth are present from both taxa an that R. latens is represented by "several hundreds" of teeth and that R. transiens is more common than the former taxon. Hence, the respective number of teeth in the taxonomic list is a rough estimate.

•Also, many teeth of Pseudocordylodon sp. 1 and Dimyloides cf. stehlini aparrently are not distinguishable from each other. The numbers given in the list are derived from the ratio of clearly assigned teeth (16:3) with a total of 52 teeth.

•Szyndlar (1994) does not specify from which of the three Egg.-Mh. localities the Bransateryx vireti vertebrae actually come. Since he states that the locality in question is of MP 30 age and only Egg.-Mh. 1 is in deposits which are not younger than Upper Oligocene, the B. vireti material has been entered here.

Reptilia
 Squamata - Boidae
Bransateryx vireti2 Hoffstetter and Rage 1972 sand boa
SMNS 56726/1, 2: 2 posterior caudal vertebrae
Mammalia
 Rodentia - Eomyidae
Rhodanomys latens Werner 1994 rodent
SMNS 44358-44362; see comments on taxonomic list
Rhodanomys transiens Hugueney 1969 rodent
SMNS 44358-44362; see comments on taxonomic list
Eomyodon mayoi Engesser 1990 rodent
SMNS 44602: 56 teeth; BSP 1983 XXII 408, 426-466: 41 teeth; EGMH-1/122-130: 8 teeth
Pseudotheridomys aff. schaubi rodent
SMNS 44363: 38 teeth; BSP 1983 XXII 404-407, 409-425: 21 teeth EGMH-1/131-135: 5 teeth
 Rodentia - Zapodidae
Plesiosminthus schaubi Viret 1926 jumping mouse
SMNS 44539, all are teeth
 Rodentia - Sciuridae
Heteroxerus lavocati squirrel
SMNS 44603, all are teeth
Heteroxerus aff. paulhiacensis squirrel
SMNS 44603: 8 teeth; BSP 1983 XXII 400-403: 4 teeth
 Rodentia - Gliridae
Peridyromys "sp. 1" Stehlin and Schaub 1951 dormouse
SMNS 44595: 354 teeth; BSP 1983 XXII 101-301, 536-548: 214 teeth; EGMH-1/55-68: 14 teeth
Microdyromys cf. monspeliensis Aguilar 1977 dormouse
SMNS 44595: 15 teeth; BSP 1983 XXII 96-100: 5 teeth; EGMH-1/109-121: 13 teeth
Bransatoglis aff. fugax Hugueney 1967 dormouse
SMNS 44597: 10 teeth; BSP 1983 XXII 74-79, 94, 95: 8 teeth; EGMH-1/44-54: 11 teeth
Bransatoglis concavidens Hugueney 1967 dormouse
SMNS 44596: 96 teeth; BSP 1983 XXII 30-73: 44 teeth; EGMH-1/9-43: 35 teeth
"Glirudinus" eggingensis Werner 1994 dormouse
SMNS 44599: 1 tooth; EGMH-1/99-108: 10 teeth
"Glirudinus" glirulus Dehm 1935 dormouse
SMNS 44600: 11 teeth; EGMH-1/76-94: 19 teeth
Suevoglis wannemacheri Werner 1994 dormouse
SMNS 44598: 18 teeth; BSP 1983 XXII 80-92: 13 teeth; EGMH-1/69-75: 7 teeth
 Rodentia - Cricetidae
Rhizospalax poirrieri rodent
SMNS 44604: 29 teeth; BSP 1983 XXII 555-567: 13 teeth
Eucricetodon longidens Hugueney 1969 rodent
SMNS 44590: 146 teeth; BSP 1983 XXII 302-348: 47 teeth
Pseudocricetodon thaleri Hugueney 1969 rodent
SMNS 44592: 8 teeth; EGMH-1/136-147: 12 teeth
Adelomyarion vireti Hugueney 1969 rodent
SMNS 44591: 171 teeth; BSP 1983 XXII 349-399: 51 teeth
 Lagomorpha - Ochotonidae
Piezodus branssatensis pika
SMNS 44606: 104 teeth; BSP 1983 XXII 549-554: 6 teeth
 Soricomorpha - Talpidae
Geotrypus sp.3 Pomel 1848 mole
SMNS 44482, all are teeth
Percymygale minor1 Ziegler 1990 mole
Myxomygale cf. engesseri3 Doukas 1986 mole
SMNS 44482, all are teeth
Hugueneya primitiva3 Hutchison 1974 mole
SMNS 44482: 23 teeth; BSP 1983 XXII 1, 3: 2 teeth
Paratalpa micheli3 Lavocat 1951 mole
SMNS 44482: 80 teeth; BSP 1983 XXII 2, 4, 6-10, 13-17: 12 teeth
 Tribosphenida - Erinaceidae
Amphechinus arvernensis4 Blainville 1839 hedgehog
SMNS 44747: 16 teeth; BSP 1983 XXII 520, 530, 531*, 532*, 535: 5 teeth; * = possibly A. robustus
 Tribosphenida - Erinaceoidae
"Pseudocordylodon "sp. 1"" = Dimyloides4, Dimyloides cf. stehlini4
"Pseudocordylodon "sp. 1"" = Dimyloides4 Hürzeler 1944 placental
SMNS 44769: 12 clearly assigned teeth; BSP 1983 XXII 517-519, 521-527: 2 clearly assigned teeth; EGMH-1/186-190: 2 clearly assigned teeth; see comments on taxonomic list
Dimyloides cf. stehlini4 Hürzeler 1944 placental
SMNS 44769: 3 clearly assigned teeth; BSP 1983 XXII 517-519, 521-527: 0 clearly assigned teeth; EGMH-1/186-190: 0 clearly assigned teeth; see comments on taxonomic list
 Tribosphenida - Herpetotheriidae
Amphiperatherium exile4 Gervais 1848 marsupial
SMNS 44660: 31 teeth; BSP 1983 XXII 467-471: 16 teeth; EGMH-1/2-6: 5 teeth