Haikou [Yuanshan Fm (formerly Qiongzhusi)] (Cambrian of China)

Also known as Chengjiang biota; Jingmacun, Kunming

Where: Yunnan, China (24.8° N, 102.6° E: paleocoordinates 35.1° N, 161.1° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Eoredlichia trilobite zone, Yuanshan Formation, Atdabanian (521.0 - 516.0 Ma)

• LITHOSTRATIGRAPHY: Formerly from the Yuanshan Mbr, Qiongzhusi Fm. Yuanshan is raised to formation rank in recent publications (Steiner 2005, 2007). Wade-Giles romanization of Qiongzhusi is Chiungchussu. AGE: Eoredlichia Zone. Refined to Atdabanian by A. Hendy (2/2009)

• group of beds-level stratigraphic resolution

Environment/lithology: marine; lithified, gray, green, yellow, carbonaceous mudstone

• "benthic sediment flows... are believed to have been the the principal method of burial" (Shu et al. 1999b)

•"presence of agglutinated quartz grains, substantially coarser than the surrounding matrix... suggests... that the animal inhabited a higher-energy, sandy sea floor and was transported" (Shu et al. 2001)

• LITHOLOGY: Dark grey-green (yellow, when weathered), finely laminated mudstones, containing organic hash in great quantity. LITHIFICATION: Lithified, on the basis of figures.

Size class: macrofossils

Preservation: adpression, soft parts

Collection methods: surface (in situ), hydrochloric,

• COLLECTOR: Multiple collectors. REPOSITORY: Not stated in text.

Primary reference: M. Steiner, M. Zhu, Y. Zhao and B.-D. Erdtmann. 2005. Lower Cambrian Burgess Shale-type fossil associations of South China. Palaeogeography, Palaeoclimatology, Palaeoecology 220:129-152 [A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 68083: authorized by John Alroy, entered by John Alroy on 02.01.2007, edited by Pete Wagner, Austin Hendy and Marcus Domansky

Creative Commons license: CC BY (attribution)

Taxonomic list

• COVERAGE: Exhaustive for all groups. NOMENCLATURE: From a combination of authoritative and review publications, with modern nomenclature and species-resolution identifications.
unclassified
  -
Sinocylindra yunnanensis
  -
Sinocylindra yunnanensis Chen and Erdtmann 1991
Fuxianospira
  -
Cambrorhytium
  -
Xianguangia
  -
Xianguangia sinica Chen and Erdtmann 1991
Hexactinellida
 Reticulosa - Protospongiidae
Saetaspongia densa Mehl and Reitner 1993 glass sponge
Demospongiae
  - Vauxiidae
Vauxia progracilenta n. sp.9 demosponge
Vauxia paraleioia9 demosponge
 Protomonaxonida - Choiidae
Choiaella radiata Rigby and Hou 1995 demosponge
Choia sp. Walcott 1920 demosponge
Allantospongia mica Rigby and Hou 1995 demosponge
 Protomonaxonida - Halichondritidae
Halichondrites sp. Walcott 1920 demosponge
 Protomonaxonida - Leptomitidae
Paraleptomitella dictyodroma Chen et al. 1989 demosponge
Leptomitus teretiusculus Chen et al. 1989 demosponge
 Protomonaxonida - Hazeliidae
Hazelia palmata Walcott 1920 demosponge
 Protomonaxonida - Hamptoniidae
Hamptonia bowerbanki Walcott 1920 demosponge
Nectocaris
  - Nectocarididae
Petalilium latus Luo and Hu 1999
Scleroctenophora
  -
Trigoides aclis Luo et al. 1999
Xidazoon stephanus
  -
Xidazoon stephanus n. gen. n. sp.7 Shu et al. 1999
Xidazoon stephanus Shu et al. 1999
Agnatha
  -
Haikouichthys ercaicunensis n. gen. n. sp.8 Luo et al. 1999 jawless fish
Haikouichthys ercaicunensis Luo et al. 1999 jawless fish
Myllokunmingia fengjiaoa n. gen. n. sp.8 Wagner 2023 jawless fish
Myllokunmingia fengjiaoa Wagner 2023 jawless fish
Cephalochordata
  -
Vetulicolida
 Vetulicolata - Didazoonidae
  - Vetulicolidae
 Bilateria - Eldoniidae
Stellostomites eumorphus Sun and Hou 1987
 Bilateria -
Cheungkongella ancestralis n. gen. n. sp.6 Shu et al. 2001
Cheungkongella ancestralis Shu et al. 2001
Cotyledion tylodes
  -
Cotyledion tylodes Luo and Hu 1999
 Lophophorata -
Hyolitha
 Hyolithida -
 Hyolithida - Sulcavitidae
Brachiopoda
 Tommotiida -
Wufengella bengtsoni n. gen. n. sp.2
Wufengella bengtsoni n. gen. n. sp.2 Guo et al. 2022
Lingulata
 Lingulida - Botsfordiidae
Diandongia pista Rong 1974
 Lingulida - Obolidae
"Lingulella chengjiangensis" = Eoglossa chengjiangensis
"Lingulella chengjiangensis" = Eoglossa chengjiangensis Jin et al. 1993
 Lingulida - Lingulellotretidae
Brachiopoda
  - Mickwitziidae
Heliomedusa orienta10 Sun and Hou 1987
Heliomedusa orienta Sun and Hou 1987
 Lophophorata -
Conicula striata Luo et al. 1999
Amiskwia
  - Amiskwidae
Cardiodictyon
  -
Cardiodictyon catenulum Hou et al. 1991
Diania cactiformis
  -
Diania cactiformis n. gen. n. sp.5
Diania cactiformis n. gen. n. sp.5 Liu et al. 2011
 Hallucishaniids -
Hallucigenia fortis1 Hou and Bergström 1995
 Bilateria -
Onychodictyon gracilis n. sp.3 Liu et al. 2008
Deuteropoda
 Fuxianhuiida - Fuxianhuiidae
Megacheira
 Leanchoiliida - Leanchoiliidae
"Leanchoilia illecebrosa" = Leanchoilia illecebrosa, "Leanchoilia asiatica" = Leanchoilia illecebrosa, "Dianchia mirabilis" = Leanchoilia illecebrosa, "Apiocephalus elegans" = Leanchoilia illecebrosa
"Leanchoilia illecebrosa" = Leanchoilia illecebrosa Hou 1987
"Leanchoilia asiatica" = Leanchoilia illecebrosa Hou 1987
"Dianchia mirabilis" = Leanchoilia illecebrosa Hou 1987
"Apiocephalus elegans" = Leanchoilia illecebrosa Hou 1987
 Fortiforcipida - Fortiforcipidae
Fortiforceps foliosa Hou and Bergström 1997
 Mollisoniida -
Urokodia aequalis Hou et al. 1989
Deuteropoda
 Isoxyida - Isoxyidae
Isoxys curvirostratus Vannier and Chen 2000
Isoxys auritus Jiang 1982
Isoxys paradoxus Hou 1987
Crustacea
 Waptiida - Waptiidae
Waptia sp.8 Walcott 1912
Deuteropoda
 Hymenostraca -
Artiopoda
 Nektaspidida -
Sinoburius lunaris Hou et al. 1991
Cindarella eucalla Chen et al. 1996
Xandarella sp. Hou et al. 1991
 Nektaspidida - Naraoiidae
Naraoia sp.8 Walcott 1912
Naraoia spinosa Zhang and Hou 1985
"Naraoia longicaudata" = Misszhouia longicaudata Zhang and Hou 1985
 Conciliterga -
Saperion glumaceum Hou et al. 1991
Kuamaia lata Hou 1987
  -
Squamacula clypeata Hou and Bergström 1997
Trilobita
 Redlichiida - Redlichiidae
Eoredlichia sp.8 Chang 1950 trilobite
"Eoredlichia intermedia" = Eoredlichia intermediata Lu 1940 trilobite
Kuanyangia pustulosa Lu 1941 trilobite
 Redlichiida - Yunnanocephalidae
Yunnanocephalus sp.8 Kobayashi 1936 trilobite
Yunnanocephalus yunnanensis Mansuy 1912 trilobite
Artiopoda
  -
Retifacies abnormalis Hou et al. 1989
Megadictyon
  -
 Radiodonta - Amplectobeluidae
 Radiodonta - Anomalocarididae
Parapeytoia yunnanensis Hou et al. 1995
 Radiodonta - Tamisiocarididae
"Anomalocaris saron" = Houcaris saron
"Anomalocaris saron" = Houcaris saron Hou et al. 1995
Jianfengia
  -
 Odaraiida - Pectocariidae
"? Odaraia eurypetala" = Pectocaris
"? Odaraia eurypetala" = Pectocaris
 Parvancorinomorpha - Parvancorinidae
Primicaris larvaformis Zhang et al. 2003
Ercaia
  -
 Bradoriida -
Tanglangia
  -
Bilateria
 Luolishaniida -
Luolishania longicruris Hou and Chen 1989
Facivermis yunnanicus Hou and Chen 1989
"Xishania longiusula" = Facivermis longiusula Hu 2002
Xenusia
 Xenusiida - Xenusiidae
Jianshanopodia decora n. gen. n. sp.4
Jianshanopodia decora n. gen. n. sp.4 Liu et al. 2006
Sicyophorus
  -
Sicyophorus rara Luo and Hu 1999
Acosmia maotiania
  -
Acosmia maotiania Chen and Zhou 1997
 Archaeopriapulida - Ottoidae
Palaeoscolecida
 Cricocosmida - Palaeoscolecidae
"Palaeoscolex sinensis" = Mafangscolex sinensis
"Palaeoscolex sinensis" = Mafangscolex sinensis Hou and Sun 1988
 Palaeoscolecidomorpha - Cricocosmiidae
 Palaeoscolecidomorpha -
? Maotianshania cylindrica Sun and Huo 1987
Corynetis brevis
  -
Corynetis brevis Luo and Hu 1999
 Selkirkiimorpha - Selkirkiidae
"Paraselkirkia sinica" = Selkirkia sinica
"Paraselkirkia sinica" = Selkirkia sinica Luo and Hu 1999
Discoides abnormis
  -
Discoides abnormis Luo et al. 1999
Sandaokania latinodosa
  -
Sandaokania latinodosa Luo and Hu 1999
Lagenula striolata
  -
Lagenula striolata Luo and Hu 1999