Haikou [Yuanshan Fm (formerly Qiongzhusi)] (Cambrian of China)

Also known as Chengjiang biota; Jingmacun, Kunming

Where: Yunnan, China (24.8° N, 102.6° E: paleocoordinates 35.1° N, 161.1° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Eoredlichia trilobite zone, Yuanshan Formation, Atdabanian (521.0 - 516.0 Ma)

• LITHOSTRATIGRAPHY: Formerly from the Yuanshan Mbr, Qiongzhusi Fm. Yuanshan is raised to formation rank in recent publications (Steiner 2005, 2007). Wade-Giles romanization of Qiongzhusi is Chiungchussu. AGE: Eoredlichia Zone. Refined to Atdabanian by A. Hendy (2/2009)

• group of beds-level stratigraphic resolution

Environment/lithology: marine; lithified, gray, green, yellow, carbonaceous mudstone

• "benthic sediment flows... are believed to have been the the principal method of burial" (Shu et al. 1999b)

•"presence of agglutinated quartz grains, substantially coarser than the surrounding matrix... suggests... that the animal inhabited a higher-energy, sandy sea floor and was transported" (Shu et al. 2001)

• LITHOLOGY: Dark grey-green (yellow, when weathered), finely laminated mudstones, containing organic hash in great quantity. LITHIFICATION: Lithified, on the basis of figures.

Size class: macrofossils

Preservation: adpression, soft parts

Collection methods: surface (in situ), hydrochloric,

• COLLECTOR: Multiple collectors. REPOSITORY: Not stated in text.

Primary reference: M. Steiner, M. Zhu, Y. Zhao and B.-D. Erdtmann. 2005. Lower Cambrian Burgess Shale-type fossil associations of South China. Palaeogeography, Palaeoclimatology, Palaeoecology 220:129-152 [A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 68083: authorized by John Alroy, entered by John Alroy on 02.01.2007, edited by Austin Hendy, Marcus Domansky and Pete Wagner

Creative Commons license: CC BY (attribution)

Taxonomic list

• COVERAGE: Exhaustive for all groups. NOMENCLATURE: From a combination of authoritative and review publications, with modern nomenclature and species-resolution identifications.
Vetulicolida
 Vetulicolata - Didazoonidae
  - Vetulicolidae
unclassified
  -
 Bilateria - Eldoniidae
Stellostomites eumorphus Sun and Hou 1987
 Bilateria -
Cheungkongella ancestralis n. gen. n. sp.3 Shu et al. 2001
Cheungkongella ancestralis Shu et al. 2001
Cephalochordata
  -
Agnatha
  -
Haikouichthys ercaicunensis n. gen. n. sp.5 Luo et al. 1999 jawless fish
Haikouichthys ercaicunensis Luo et al. 1999 jawless fish
Myllokunmingia fengjiaoa n. gen. n. sp.5 Wagner 2023 jawless fish
Myllokunmingia fengjiaoa Wagner 2023 jawless fish
Amiskwia
  - Amiskwidae
 Lophophorata -
Hyolitha
 Hyolithida - Sulcavitidae
 Hyolithida -
 Lophophorata -
Conicula striata Luo et al. 1999
Lingulata
 Discinida - Disciniidae
Heliomedusa orienta7 Sun and Hou 1987
Heliomedusa orienta Sun and Hou 1987
 Lingulida - Botsfordiidae
Diandongia pista Rong 1974
 Lingulida - Lingulellotretidae
 Lingulida - Obolidae
"Lingulella chengjiangensis" = Eoglossa chengjiangensis
"Lingulella chengjiangensis" = Eoglossa chengjiangensis Jin et al. 1993
Cotyledion tylodes
  -
Cotyledion tylodes Luo and Hu 1999
Megadictyon
  -
 Mollisoniida -
Urokodia aequalis Hou et al. 1989
Deuteropoda
 Isoxyida - Isoxyidae
Isoxys auritus Jiang 1982
Isoxys paradoxus Hou 1987
Isoxys curvirostratus Vannier and Chen 2000
 Fuxianhuiida - Fuxianhuiidae
 Hymenostraca -
Crustacea
 Waptiida - Waptiidae
Waptia sp.5 Walcott 1912
Megacheira
 Leanchoiliida - Leanchoiliidae
"Leanchoilia illecebrosa" = Leanchoilia illecebrosa, "Leanchoilia asiatica" = Leanchoilia illecebrosa, "Apiocephalus elegans" = Leanchoilia illecebrosa, "Dianchia mirabilis" = Leanchoilia illecebrosa
"Leanchoilia illecebrosa" = Leanchoilia illecebrosa Hou 1987
"Leanchoilia asiatica" = Leanchoilia illecebrosa Hou 1987
"Apiocephalus elegans" = Leanchoilia illecebrosa Hou 1987
"Dianchia mirabilis" = Leanchoilia illecebrosa Hou 1987
 Fortiforcipida - Fortiforcipidae
Fortiforceps foliosa Hou and Bergström 1997
 Radiodonta - Amplectobeluidae
 Radiodonta - Anomalocarididae
Anomalocaris saron Hou et al. 1995
Parapeytoia yunnanensis Hou et al. 1995
Artiopoda
  -
Squamacula clypeata Hou and Bergström 1997
 Nektaspidida - Naraoiidae
Naraoia sp.5 Walcott 1912
Naraoia spinosa Zhang and Hou 1985
Naraoia longicaudata Zhang and Hou 1985
 Nektaspidida -
Sinoburius lunaris Hou et al. 1991
Xandarella sp. Hou et al. 1991
Cindarella eucalla Chen et al. 1996
  -
Retifacies abnormalis Hou et al. 1989
Trilobita
 Redlichiida - Redlichiidae
Eoredlichia sp.5 Chang 1950 trilobite
"Eoredlichia intermedia" = Eoredlichia intermediata Lu 1940 trilobite
Kuanyangia pustulosa Lu 1941 trilobite
 Redlichiida - Yunnanocephalidae
Yunnanocephalus sp.5 Kobayashi 1936 trilobite
Yunnanocephalus yunnanensis Mansuy 1912 trilobite
Artiopoda
 Conciliterga -
Saperion glumaceum Hou et al. 1991
Kuamaia lata Hou 1987
 Parvancorinomorpha - Parvancorinidae
Primicaris larvaformis Zhang et al. 2003
 Bradoriida -
Tanglangia
  -
 Odaraiida - Pectocariidae
"? Odaraia eurypetala" = Pectocaris
"? Odaraia eurypetala" = Pectocaris
Ercaia
  -
Jianfengia
  -
 Bilateria -
Diania cactiformis
  -
Diania cactiformis n. gen. n. sp.2
Diania cactiformis n. gen. n. sp.2 Liu et al. 2011
 Hallucishaniids -
Hallucigenia fortis1 Hou and Bergström 1995
Cardiodictyon
  -
Cardiodictyon catenulum Hou et al. 1991
Bilateria
 Luolishaniida -
"Xishania longiusula" = Facivermis longiusula Hu 2002
Facivermis yunnanicus Hou and Chen 1989
Luolishania longicruris Hou and Chen 1989
Corynetis brevis
  -
Corynetis brevis Luo and Hu 1999
Acosmia maotiania
  -
Acosmia maotiania Chen and Zhou 1997
Paraselkirkia
  -
 Archaeopriapulida - Ottoidae
 Palaeoscolecidomorpha -
? Maotianshania cylindrica Sun and Huo 1987
Palaeoscolecida
 Cricocosmida - Palaeoscolecidae
"Palaeoscolex sinensis" = Mafangscolex sinensis
"Palaeoscolex sinensis" = Mafangscolex sinensis Hou and Sun 1988
 Palaeoscolecidomorpha - Cricocosmiidae
Sicyophorus
  -
Sicyophorus rara Luo and Hu 1999
Scleroctenophora
  -
Trigoides aclis Luo et al. 1999
Nectocaris
  - Nectocarididae
Petalilium latus Luo and Hu 1999
Xidazoon stephanus
  -
Xidazoon stephanus n. gen. n. sp.4 Shu et al. 1999
Xidazoon stephanus Shu et al. 1999
Cambrorhytium
  -
Xianguangia
  -
Xianguangia sinica Chen and Erdtmann 1991
Hexactinellida
 Reticulosa - Protospongiidae
Saetaspongia densa Mehl and Reitner 1993 glass sponge
Demospongiae
  - Vauxiidae
Vauxia progracilenta n. sp.6 demosponge
Vauxia paraleioia6 demosponge
 Protomonaxonida - Hazeliidae
Hazelia palmata Walcott 1920 demosponge
 Protomonaxonida - Hamptoniidae
Hamptonia bowerbanki Walcott 1920 demosponge
 Protomonaxonida - Leptomitidae
Leptomitus teretiusculus Chen et al. 1989 demosponge
Paraleptomitella dictyodroma Chen et al. 1989 demosponge
 Protomonaxonida - Choiidae
Allantospongia mica Rigby and Hou 1995 demosponge
Choia sp. Walcott 1920 demosponge
Choiaella radiata Rigby and Hou 1995 demosponge
 Protomonaxonida - Halichondritidae
Halichondrites sp. Walcott 1920 demosponge
Sinocylindra yunnanensis
  -
Sinocylindra yunnanensis Chen and Erdtmann 1991
Fuxianospira
  -
Sandaokania latinodosa
  -
Sandaokania latinodosa Luo and Hu 1999
Lagenula striolata
  -
Lagenula striolata Luo and Hu 1999
Discoides abnormis
  -
Discoides abnormis Luo et al. 1999