Issel (Eocene to of France)

Where: Aude, France (43.4° N, 2.0° E: paleocoordinates 39.6° N, 0.8° W)

• coordinate estimated from map

• small collection-level geographic resolution

When: Sables du Castrais Formation, Ypresian to Ypresian (56.0 - 38.0 Ma)

• Within the Sables du Castrais Formation at Issel, several fossiliferous levels have produced vertebrate remains with different ages spanning the upper Lutetian–upper Bartonian interval (Marandat, 1987). Contrary to previous propositions of a Bartonian age, Godinot et al. (2018) suggested that the locality of Issel could be either Ypresian–Lutetian or Lutetian, and view the holotype mandible of Dentaneosuchus crassiproratus from Issel as coming from a level situated below the base of the Lutetian (Martin et al. 2023)

• group of beds-level stratigraphic resolution

Environment/lithology: "channel"; muddy sandstone

Size class: macrofossils

Collected by Mr Henri de Sévérac in 1873

Primary reference: M. Richard. 1946. Contribution a l'étude du basin d'Aquitaine, Les Gisements de Mammiferes Tertiares. Memoires de la Societe Geologique de France, Nouvelle Serie 24(52):1-380 [M. Uhen/M. Uhen/M. Uhen]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 78233: authorized by Mark Uhen, entered by Mark Uhen on 31.01.2008, edited by Philip Mannion, Patricia Holroyd and David Nicholson

Creative Commons license: CC BY (attribution)

Taxonomic list

Mammalia
 Creodonta - Hyaenodontidae
Hyaenodon sp. Laizer and Parieu 1838 creodont
 Perissodactyla - Palaeotheriidae
Propalaeotherium sp., "Propalaeotherium parvulum" = Eurohippus parvulus, "Propalaeotherium isselanum" = Palaeotherium isselanum
Propalaeotherium sp. Gervais odd-toed ungulate
"Propalaeotherium parvulum" = Eurohippus parvulus Laurillard 1849 odd-toed ungulate
"Propalaeotherium isselanum" = Palaeotherium isselanum Cuvier 1824 odd-toed ungulate
 Perissodactyla - Lophiodontidae
Lophiodon sp. Cuvier 1822 odd-toed ungulate
Lophiodon isselense Fischer 1829 odd-toed ungulate
Paralophiodon isselense2 odd-toed ungulate
Lophiaspis occitanicus Deperet 1910 odd-toed ungulate
Reptilia
 Testudines - Podocnemididae
Neochelys sp.3 Bergounioux 1954 sideneck turtle
Listed by Broin 1977 as Neochelys eocaenica
 Testudines - Trionychidae
Trionyx sp.1 Geoffroy Saint-Hilaire 1809 softshell turtle
 Loricata -
Dentaneosuchus crassiproratus n. gen.4 Astre 1931 crocodilian
MHNT unnumbered - type (anterior portion of mandible)
 Crocodylia -
Kentisuchus astrei n. sp.2, "Crocodylus depressifrons" = Asiatosuchus depressifrons2
Kentisuchus astrei n. sp.2 Jouve 2016 crocodilian
Unprepared skull housed in the Museum d'Histoire Naturelle de Toulouse (assigned to A. glareae by Astre 1931)
"Crocodylus depressifrons" = Asiatosuchus depressifrons2 Blainville 1855 crocodilian
ISS 1-3 (cranial and mandibular material)