Woodring Loc. 139c - eastern area [lower Gatun Fm] (Miocene of Panama)

Also known as USGS #22018

Where: Panama (9.3° N, 79.8° W: paleocoordinates 9.1° N, 78.5° W)

• coordinate stated in text

• outcrop-level geographic resolution

When: lower Gatun Member (Gatun Formation), Tortonian (11.6 - 7.2 Ma)

• Lower part of the Gatun Formation. The Gatun Formation is sibdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Late Miocene, on the basis of microfossil biostratigraphy (see Collins and Coates, 1999), refined by enterer to Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 10.9 Ma (Hendy, unpublished data).

• bed-level stratigraphic resolution

Environment/lithology: marine; unlithified, fine-grained, silty sandstone

• No specific environmental data presented.
• LITHOLOGY: Soft, silty fine-grained sandstone.

Size class: macrofossils

Reposited in the USNM

Collection methods: surface (in situ),

• COLLECTOR: Not stated in text. REPOSITORY: USNM.

Primary reference: W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 93160: authorized by Austin Hendy, entered by Austin Hendy on 06.01.2010

Creative Commons license: CC BY (attribution)

Taxonomic list

• COVERAGE: Exhaustive for gastropods and bivalves. NOMENCLATURE: Authoritative publications, with modern nomenclature, and species-resolution identifications.
Bivalvia
 Nuculanida - Nuculanidae
Saccella acrita6 Dall 1908 pointed nut clam
SUBSPECIES: Saccella acrita epacra
 Nuculida - Nuculidae
Nucula (Nucula) cf. cahuitensis6 Olsson 1922 nut clam
 Carditida - Crassatellidae
Crassinella martinicensis d'Orbigny 1853 clam
 Solenida - Solenidae
Solen (Solena) obliquus Spengler 1794 clam
SUBSPECIES: Solen (Solena) obliquus rudis
 Pholadida - Corbulidae
Caryocorbula oropendula Olsson 1922 clam
SUBSPECIES: Caryocorbula oropendula stena
Varicorbula disparilis d'Orbigny 1853 clam
 Cardiida - Cardiidae
"Trigoniocardia (Apiocardia) aminensis" = Trigoniocardia (Trigoniocardia) aminensis Dall 1900 cockle
 Cardiida - Semelidae
Ervilia valhosierr Gardner 1928 clam
 Cardiida - Tellinidae
Tellina (Eurytellina) angulosa Gmelin 1791 tellin clam
SUBSPECIES: Tellina (Eurytellina) angulosa costaricana
Psammacoma gatunensis Toula 1909 tellin clam
Tellina (Merisca) aequistriata Say 1824 tellin clam
 Cardiida - Kelliellidae
Alveinus rotundus Dall 1900 clam
 Cardiida - Chamidae
Chama berjadinensis Hodson 1927 jewel box
 Cardiida - Veneridae
Chione (Chionopsis) tegulum Brown and Pilsbry 1911 venus clam
Lirophora (Lirophora) falconensis Hodson et al. 1927 venus clam
Pitar (Hyphantosoma) centangulatus Brown and Pilsbry 1911 venus clam
"Agriopoma (Pitarella) gatunensis" = Callocardia (Agriopoma) gatunensis Dall 1903 venus clam
"Macrocallista (Paradione) maculata" = Macrocallista maculata Linnaeus 1758 venus clam
Cyclinella cyclica Guppy 1866 venus clam
Clementia (Clementia) dariena Conrad 1855 venus clam
SUBSPECIES: Clementia (Clementia) dariena dariena
Gouldia costaricensis Olsson 1922 venus clam
 Lucinida - Lucinidae
Bellucina actinus Dall 1903 clam
 Ostreida - Gryphaeidae
Hyotissa haitensis Sowerby 1850 oyster
 Ostreida - Pteriidae
Pteria inornata6 Gabb 1873 pearl oyster
 Pectinida - Pectinoidae
 Arcida - Arcidae
Anadara (Rasia) dariensis6 Brown and Pilsbry 1911 ark
Anadara (Rasia) fissicosta6 Spieker 1922 ark
 Mytilida - Crenellidae
"Crenella divaricata" = Crenella decussata6
"Crenella divaricata" = Crenella decussata6 Montagu 1808 clam
Scaphopoda
 Gadilida - Gadilidae
Cadulus (Platyschides) epetrion6, "Cadulus (Gadilopsis) dentalinus" = Gadilopsis6
"Cadulus (Gadilopsis) dentalinus" = Gadilopsis6 tusk shell
 Dentaliida - Dentaliidae
Gastropoda
 Opisthobranchia -
Roxania chipolana5 Dall 1895 snail
 Opisthobranchia - Retusidae
Sulcoretusa sulcata5 d'Orbigny 1841 snail
SUBSPECIES: Sulcoretusa sulcata lipara
 Opisthobranchia - Cylichnidae
Cylichnella atacata5 Woodring 1928 snail
 Heterostropha - Ringiculidae
 Heterostropha - Acteonidae
Rictaxis oryza5, "Acteon (Acteon) punctostriatus" = Japonactaeon punctostriata5
Rictaxis oryza5 Gabb 1872 snail
"Acteon (Acteon) punctostriatus" = Japonactaeon punctostriata5 Adams 1840 snail
 Vanikoroidea - Eulimidae
Balcis (Balcis) lipara5 Woodring 1970 snail
Balcis (Balcis) cetia5 Woodring 1970 snail
Balcis (Balcis) jacululum5 Maury 1917 snail
Eulima nobilis5 Guppy 1896 snail
 Neogastropoda - Olividae
Olivella (Niteoliva) terryi4, "Oliva (Oliva) gatunensis" = Oliva gatunensis4
Olivella (Niteoliva) terryi4 Olsson 1956 olive snail
"Oliva (Oliva) gatunensis" = Oliva gatunensis4 Toula 1908 olive snail
 Neogastropoda - Ancillariidae
Ancilla (Eburna) pinguis4 Guppy 1873 snail
 Neogastropoda - Marginellidae
"Prunum (Microspira) gatunense" = Prunum gatunense5
"Prunum (Microspira) gatunense" = Prunum gatunense5 Brown and Pilsbry 1911 margin shell
 Neogastropoda - Cancellariidae
"Cancellaria (Euclia) codazzii" = Euclia codazzii5, Bivetiella dariena1, Cancellaria acalypta5, "Cancellaria (Narona) barystoma" = Narona barystoma5, "Trigonostoma cf. scalatella" = Axelella scalatella5
"Cancellaria (Euclia) codazzii" = Euclia codazzii5 Anderson 1929 snail
Bivetiella dariena1 Toula 1908 snail
SUBSPECIES: Cancellaria (Cancellaria) epistomifera dariensis
Cancellaria acalypta5 Woodring 1970 snail
"Cancellaria (Narona) barystoma" = Narona barystoma5 Woodring 1970 snail
"Trigonostoma cf. scalatella" = Axelella scalatella5 Guppy 1873 snail
 Neogastropoda - Turbinellidae
"Xancus dodonaius" = Turbinella6, "Xancus validus" = Turbinella valida4
"Xancus dodonaius" = Turbinella6 snail
SUBSPECIES: Xancus dodonaius praelaeviagtus
"Xancus validus" = Turbinella valida4 Sowerby 1850 snail
SUBSPECIES: Xancus validus falconensis
 Neogastropoda - Nassariidae
Calophos ectyphus4 Woodring 1964 snail
 Neogastropoda - Buccinidae
Strombinophos mimicus4, "Antillophos (Antillophos) candei" = Antillophos candeanus4
Strombinophos mimicus4 Woodring 1964 true whelk
"Antillophos (Antillophos) candei" = Antillophos candeanus4 d'Orbigny 1842 true whelk
 Neogastropoda - Fasciolariidae
"Fasciolaria gorgasiana" = Pleuroploca gorgasiana4
"Fasciolaria gorgasiana" = Pleuroploca gorgasiana4 Brown and Pilsbry 1913 snail
 Neogastropoda - Columbellidae
Strombina (Strombina) lessepsiana4 Brown and Pilsbry 1911 snail
Anachis (Costoanachis) mira4 Dall 1896 snail
SUBSPECIES: Anachis (Costoanachis) mira mira
 Neogastropoda - Drilliidae
Clathrodrillia gatunensis5 Toula 1909 snail
"Crassispira (Hindsiclava) consors" = Hindsiclava consors5 Sowerby 1850 snail
Agladrillia (Eumetadrillia) isthmica5 Brown and Pilsbry 1911 snail
Agladrillia (Agladrillia) enneacyma5 Brown and Pilsbry 1913 snail
Agladrillia (Agladrillia) acaria5 Mansfield 1925 snail
SUBSPECIES: Agladrillia (Agladrillia) acaria ectypha
 Neogastropoda - Conidae
"Conus multiliratus" = Conasprelloides multiliratus5 Olsson 1922 cone shell
Conus burckhardti5 Bose 1906 cone shell
Conus aemulator5 Brown and Pilsbry 1911 cone shell
 Neogastropoda - Mangeliidae
Kurtziella (Kurtziella) pagella5 Woodring 1970 snail
Dolostoma anorhepes5 Woodring 1970 snail
 Neogastropoda - Borsoniidae
Microdrillia trina5 Mansfield 1925 snail
 Neogastropoda - Clathurellidae
Nannodiella rintriada5 Mansfield 1925 snail
 Neogastropoda - Turridae
"Pleurofusia (Cruziturricula) fusinus" = Cruziturricula fusinus5, Polystira sp.2
"Pleurofusia (Cruziturricula) fusinus" = Cruziturricula fusinus5 Brown and Pilsbry 1911 turrid
Polystira sp.2 Woodring 1928 turrid
 Neogastropoda - Terebridae
Strioterebrum spiriferum5 Dall 1895 auger snail
Strioterebrum oresignum5 Olsson 1964 auger snail
Strioterebrum indocayapum5 Olsson 1964 auger snail
Terebra (Panaterebra) cucurrupiensis5 Oinomikado 1939 auger snail
Terebra (Oreoterebra) subsulcifera5 Brown and Pilsbry 1911 auger snail
Terebra (Oreoterebra) isaacpetiti5 Maury 1925 auger snail
 Sorbeoconcha - Vermetidae
Petaloconchus sculpturatus5 Lea 1846 worm shell
 Pseudomelanoidea - Pseudomelaniidae
"Carinodrillia (Carinodrillia) cf. elocata" = Carinodrillia elocata5 Pilsbry and Johnson 1922 snail