Full reference: G. Bechly. 1996. Morphologische Untersuchungen am Flügelgeäder der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Berücksichtigung der phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special-Volume 2:1-402
Parent taxon: Anisoptera according to G. Bechly 2007
See also Bechly et al. 1998 and Ren et al. 2003
Sister taxa: Aeschna eudore, Aeschna parkinsoni, Aeschnidioidea, Cercus, Cercus clavus, Cordulia scheuchzeri, Cratopetaliidae, Cymatophlebia mongolica, Dissurus, Dissurus liauyuanensis, Dissurus qinquanensis, Epigomphidae, Hageniidae, Henrotayiidae, Ictinogomphus fur, Ixtahua, Juragomphidae, Kazakhophlebiella, Libanoaeshnidae, Libellula calypso, Libellula doris, Libellula eurynome, Libellula melobasis, Libellula perse, Libellula thetis, Libellula thoe, Libelluloidea, Magnathemidae, Megaphlebiidae, Miopetalura orientalis, Necrogomphus, Nothomacromiidae, Petalurida, Pritykiniella, Proterogomphidae, Stenogomphus scudderi
Subtaxa: Aeshnoptera Exophytica
Ecology: fast-moving volant carnivore-insectivore