Full reference: G. Bechly. 1996. Morphologische Untersuchungen am Flügelgeäder der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Berücksichtigung der phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special-Volume 2:1-402
Parent taxon: Odonata according to G. Bechly 2007
See also Bechly 2005, Bechly et al. 2003 and Petrulevičius and Nel 2007
Sister taxa: Anisozygoptera, Hageniella, Hageniella problematica, Haidilaozhenidae, Idionychidae, Kobdoneura, Liassophlebia clavigaster, Liassophlebia hopei, Libellula decapitata, Libellulina, Mesomantidiidae, Mesonetopsis, Orthaeschnites, Oryctothemis, Palaeophlebia, Peraphlebia, Proaeschna, Pseudoeuphaea, Pseudoeuphaea areolata, Pseudoeuphaea falsificata, Pseudoeuphaea filosa, Pseudoeuphaea obscura, Sogdothemis, Syrrhoe, Triassophlebia, Zygoptera
Subtaxa: Euepiproctophora Frenguelliidae Isophlebioptera
Ecology: fast-moving volant carnivore-insectivore