Protainophthalmus asperulus Heer 1856

Insecta - Coleoptera - Curculionidae

Alternative combination: Cleonus asperulus

Synonyms: Brachyderes aquisextanus Oustalet 1874, Brachyderes longipes Oustalet 1874, Cleonus marcelli Oustalet 1874

Full reference: O. Heer. 1856. Ueber die fossilen Insekten von Aix in der Provence. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich 1:1-40

Belongs to Protainophthalmus according to A. A. Legalov 2015

See also Förster 1891, Heer 1856, Oustalet 1874, Theobald 1937, Yunakov and Kirejtshuk 2011 and Zherikhin 1992

Sister taxa: Protainophthalmus margarus, Protainophthalmus punctulatus, Protainophthalmus regularis, Protainophthalmus rugicollis, Protainophthalmus thaisi, Protainophthalmus tuberculatus

Type specimens:

  • Protainophthalmus asperulus: Its type locality is Aix-en-Provence (Musee Blanchet Coll), which is in a Chattian lacustrine - large limestone in the Niveau du gypse d'Aix Formation of France
  • Brachyderes aquisextanus:
  • Brachyderes longipes:
  • Cleonus marcelli:


• Miocene of Italy (1 collection)

• Oligocene of France (5)

Total: 6 collections each including a single occurrence

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Specimen images are retrieved through the ePANDDA API.

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