Insecta - Coleoptera - Scarabaeidae
Full reference: W. E. Leach. 1815. Entomology. The Edinburgh Encyclopaedia 9:57-172
Parent taxon: Aphodiini according to R. P. Webster 2016
See also Bouchard et al. 2011
Sister taxa: Agoliinus, Caligodorus, Dialytes, Dialytodius, Pardalosus, Planolinoides
Subtaxa: Aphodius Diapterna Heptaulacus
Ecology:
Distribution:
• Quaternary of Greenland (1 collection), Japan (6), the Russian Federation (28), Ukraine (1), the United Kingdom (26), United States (18: Colorado, Pennsylvania, Washington)
• Pliocene of Germany (1)
• Miocene of Germany (3), Italy (1), Japan (1)
• Oligocene of France (1), Germany (2)
• Eocene of France (1), the United Kingdom (1), United States (7: Colorado)
• Paleocene of France (1)
• Cretaceous of the Russian Federation (1)
Total: 100 collections including 138 occurrences