Full reference: A. G. Ponomarenko. 1969. Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]. Trudy Akademiya Nauk SSSR 125:1-240
Type specimen: PIN 2555/1666, an exoskeleton. Its type locality is Madygen, PIN collection 2555, Dzhailoucho area, which is in a Carnian lacustrine - small siliciclastic in the Madygen Formation of Kyrgyzstan.
• Triassic of Kyrgyzstan (2 collections)
Total: 2 collections each including a single occurrence
Specimen images are retrieved through the ePANDDA API.
Click image to enlarge. Click to access iDigBio record.