Full reference: A. G. Ponomarenko. 1969. Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]. Trudy Akademiya Nauk SSSR 125:1-240
Sister taxa: none
Type specimen: PIN 371/2351, an exoskeleton. Its type locality is Ak-Bulak-Say, Sogjuta, Issyk-Kul (PIN Collection 371), which is in a Hettangian/Sinemurian lacustrine - small siliciclastic in the Dzhil Formation of Kyrgyzstan.
Distribution: found only at Ak-Bulak-Say, Sogjuta, Issyk-Kul (PIN Collection 371)
Specimen images are retrieved through the ePANDDA API.
Click image to enlarge. Click to access iDigBio record.