Insecta - Coleoptera - Carabidae

Alternative combination: Shanwangicarabus brunneus

Belongs to Calosoma according to J. Zhang et al. 1994

See also Zhang 1989

Sister taxa: Calosoma (Campalita), Calosoma agassizi, Calosoma calvini, Calosoma caraboides, Calosoma catenulatum, Calosoma ceresti, Calosoma cockerelli, Calosoma emmonsi, Calosoma escheri, Calosoma escrobiculatum, Calosoma fernquisti, Calosoma grasui, Calosoma heeri, Calosoma inquisitor, Calosoma jaccardi, Calosoma kimi, Calosoma maderae, Calosoma nauckianum, Calosoma semilaeve, Calosoma sycophanta

Type specimen: 750138, an exoskeleton

Ecology:

Distribution: found only at Shanwang, Linqu County, collection 5 (Miocene of China)

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Specimen images are retrieved through the ePANDDA API.