Insecta - Coleoptera - Hydrophilidae
Full reference: J. Zhang, B. Sun, and X. Zhang. 1994. Miocene insects and spiders from Shanwang, Shandong 1-298
Parent taxon: Hydrophilidae according to J. Zhang et al. 1994
Sister taxa: Acidocerinae, Alegorius, Aposphinctus, Chaetarthriinae, Creniphilites, Cretosperchus, Enochrinae, Escheria, Escheria bella, Escheria ovalis, Georissinae, Hydrocharis, Hydrochinae, Hydrophilinae, Mesosperchus schultzi, Orphnospercheus, Paraspercheus, Philhydrus, Prospercheus, Protochares, Regimbartia, Rygmodinae, Spercheinae, Sphaeridiinae, Sphaeridium melanarium, Sternolophus
Subtaxa: Penitus gemellus
Type: Penitus gemellus
Ecology:
Distribution: found only at Shanwang, Linqu County, collection SK (Shanwang Fossil Protection Post coll.) (Miocene of China)
Specimen images are retrieved through the ePANDDA API.
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