Insecta - Coleoptera - Coptoclavidae
Full reference: C. Soriano, A. G. Ponomarenko, and X. Delclòs. 2007. Coptoclavid beetles (Coleoptera: Adephaga) from the Lower Cretaceous of Spain: A new feeding strategy in beetles. Palaeontology 50(2):525-536
Parent taxon: Coptoclavidae according to A. G. Ponomarenko and D. N. Fedorenko 2022
See also Cai et al. 2022, Ponomarenko 2014, Ponomarenko 2015 and Soriano et al. 2007
Sister taxa: Agrascapha, Amblycephalonius, Charonoscaphinae, Coptoclavia, Coptoclavinae, Euroscapha, Hispanoclavinae, Timarchopsinae
Subtaxa: Coptoclavella Coptoclavisca Holcoptera
Ecology:
Distribution:
• Cretaceous of China (1 collection), Mongolia (6), the Russian Federation (2), Spain (1), the United Kingdom (1)
• Jurassic to Cretaceous of the Russian Federation (1)
• Jurassic of Mongolia (1), the Russian Federation (2), the United Kingdom (17), United States (3: Connecticut, Massachusetts)
• Triassic to Jurassic of the United Kingdom (1)
• Triassic of Germany (4), Italy (1), Ukraine (1), the United Kingdom (7), United States (2: North Carolina)
Total: 51 collections including 59 occurrences