Insecta - Coleoptera - Scarabaeidae

Full reference: E. L. Geoffroy. 1762. Histoire Abregée des Insectes qui se Trouvent aux Environs de Paris, Tome Premier 1-523

Parent taxon: Copris according to S. Tarasov et al. 2016

See also Kabakov 1988

Sister taxa: Copris druidum, Copris incertus, Copris leakeyorum, Copris lunaris, Copris pristinus

Subtaxa: Copris (Copris) kartlinus

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Ecology:

Distribution: found only at Kisatibi village (Pliocene of Georgia)

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