Calcitoryctes Krell 2011 (rhinoceras beetle)

Insecta - Coleoptera - Scarabaeidae

Full reference: F. T. Krell. 2011. Scarabaeidae, in Beetles (Insecta: Coleoptera). Paleontology and Geology of Laetoli: Human Evolution in Context. Volume 2: Fossil Hominins and the Associated Fauna 535-548

Parent taxon: Dynastinae according to F. T. Krell 2011

Sister taxa: Cyclocephala, Ligyrus, Oryctini, Oryctoantiquus, Pentodon proserpinae, Pentodontini

Subtaxa: Calcitoryctes magnificus

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Type: Calcitoryctes magnificus

Ecology:

Distribution: found only at Laetoli Locality 2 (Pliocene of Tanzania)

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