Insecta - Coleoptera - Scarabaeidae
Parent taxon: Aphodius according to Mulsant and Rey 1870
Sister taxa: Aphodius (Acrossus), Aphodius (Nialus), Aphodius (Plagiogonus), Aphodius aboriginalis, Aphodius anteactus, Aphodius boryslavicus, Aphodius bosniaskii, Aphodius brevipennis, Aphodius carpetanus, Aphodius charauxi, Aphodius conspurcatus, Aphodius cretaceous, Aphodius distinctus, Aphodius fimitarius, Aphodius florissantensis, Aphodius fossor, Aphodius granarioides, Aphodius granarius, Aphodius helvolus, Aphodius holdereri, Aphodius inundatus, Aphodius krantzi, Aphodius laminicola, Aphodius mediaevus, Aphodius meyeri, Aphodius plagiatus, Aphodius praeemptor, Aphodius precursor, Aphodius rhinocerontis, Aphodius rufipes, Aphodius ruthenus, Aphodius schlickumi, Aphodius senex, Aphodius shoshonis, Aphodius subater, Aphodius theobaldi, Aphodius vectis
Subtaxa: none
Ecology:
Distribution: found only at Chernyi Yar (Quaternary of Russian Federation)