Insecta - Coleoptera - Scarabaeidae

Full reference: I. Fujiyama. 1968. A Miocene fossil of tropical dung beetle from Noto, Japan (Tertiary insect fauna of Japan, 2). Bulletin of the National Science Museum 11:201-210

Belongs to Heliocopris according to S. Tarasov et al. 2016

See also Fujiyama 1968

Sister taxa: none

Type specimen: Kyoto University JC 500528, an exoskeleton. Its type locality is Takaya, Suzu-shi, Noto Peninsula, which is in a Miocene terrestrial siliciclastic in the Yanagida Formation of Japan.

Ecology:

Distribution: found only at Takaya, Suzu-shi, Noto Peninsula

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Specimen images are retrieved through the ePANDDA API.