Mammalia - Artiodactyla - Bovidae
Parent taxon: Caprinae according to A. W. Gentry 2010
See also Grubb 2001
Sister taxa: Aragoral, Benicerus, Damalavus, Myotragus, Naemorhedini, Rupicaprini
Subtaxa: Makapania Nitidarcus Speleotherium
Ecology: ground dwelling grazer-browser
Distribution:
• Quaternary of Canada (2: Yukon collections), Ethiopia (3), France (1), Italy (1), the Russian Federation (1), South Africa (2), Spain (1), United States (5: California, New Mexico, Tennessee, Virginia)
• Villafranchian of Greece (1), Spain (1)
• Pliocene to Pleistocene of China (1), South Africa (1)
• Pliocene of Kenya (1), South Africa (3)
• Miocene to Pliocene of South Africa (1)
• Miocene to Pleistocene of Spain (1)
• Miocene of Iraq (1)
Total: 27 collections including 29 occurrences