Insecta - Coleoptera - Scarabaeidae

Parent taxon: Onthophagini according to T. K. Philips 2016

See also Zhang 1989

Sister taxon: Onthophagus

Subtaxa: none

Ecology:

Distribution: found only at Shanwang, Linqu County, collection s (Shandong Provincial Museum) (Miocene of China)

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Specimen images are retrieved through the ePANDDA API.