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Jormungandr walhallaensis

Reptilia - Mosasauridae

Taxonomy
Jormungandr walhallaensis was named by Zietlow et al. (2023). Its type specimen is NDGS 10838, a partial skeleton (A nearly complete skull, complete mandibles, all seven cervical vertebrae, 11 disarticulated ribs, and five articulated anterior dorsal vertebrae), and it is a 3D body fossil. Its type locality is Cavalier County, which is in a Campanian offshore shale in the Pierre Shale Formation of North Dakota. It is the type species of Jormungandr.

Synonymy list
YearName and author
2023Jormungandr walhallaensis Zietlow et al. pp. 6-8

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
RankNameAuthor
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Squamata()
familyMosasauridae
subfamilyMosasaurinaeGervais 1853
genusJormungandr
specieswalhallaensis

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Jormungandr walhallaensis Zietlow et al. 2023
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Diagnosis
ReferenceDiagnosis
A. R. Zietlow et al. 2023Mosasaurine mosasaur possessing the following suite of characters: edentulous
premaxillary rostrum of intermediate length (e.g., longer than one tooth position but shorter than two tooth positions) and blunt in lateral view but pointed in dorsoventral views; premaxilla internarial bar T-shaped in cross section along its entire length; presence of premaxilla dorsal keel; 15 maxillary teeth; frontal of intermediate width (i.e., between 1.5 and 2 times longer than wide) and possesses a low but distinct dorsal keel; presence of postorbitofrontal transverse ridge; jugal angle slightly
obtuse; quadrate stapedial pit a narrow (1.8× taller than wide) oval; quadrate posteroventral ascending rim a high, triangular crest; groove present in anterolateral portion of the quadrate alar rim; quadrate mandibular condyle distinctly convex in all views; 16 pterygoid teeth; 16 dentary teeth; dentary anterior projection longer than one tooth position); splenial–angular articular surfaces grooved; zygosphenes and zygantra present on cervical vertebrae; hypapophyses present on second through seventh cervicals; and hypapophyseal peduncle lacking on cervical seven. It is also distinguished from all other known mosasaurids by: a quadrate stapedial pit set within a sulcus that is bounded ventrally by a ridge; both maxillae and dentaries that are twice as wide as the tooth bases (as opposed to only slightly wider in other taxa); and prefrontal-postorbitofrontal contact that is prevented by a thin ridge of the frontal.