PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Marsileaceae

Polypodiopsida - Marsileaceae

Taxonomy

Marsileaceae was named by Mirbel (1802). It is extant.

It was assigned to Filicales by Knowlton (1919); to Polypodiopsida by Vera et al. (2022), Perez Loinaze et al. (2023); and to Salviniales by Jud (2011), Hermsen et al. (2013), I (2016), Vallati et al. (2020), De Benedetti et al. (2024).

Subtaxa

Collarisporites, Crybelosporites, Cystorrhiza, Gabonisporis, Lugiomarsiglia, Marsilea, Marsileaceaephyllum, Mirasolita, Molaspora, Pillularia, Regnellidium, Rhizocarpites, Rodeites

Synonyms

Cystorrhiza was named by Massalongo (1859). It is not extant.

It was considered a nomen nudum by Andrews (1970).

Synonymy list

Year	Name and author
1802	Marsileaceae Mirbel
1859	Cystorrhiza Massalongo
1919	Marsileaceae Knowlton
2011	Marsileaceae Jud
2013	Marsileaceae Hermsen et al.
2016	Marsileaceae I
2020	Marsileaceae Vallati et al. p. 5
2022	Marsileaceae Vera et al.
2023	Marsileaceae Perez Loinaze et al.
2024	Marsileaceae De Benedetti et al. p. 7

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Rank	Name	Author
kingdom	Plantae
phylum	Pteridophyta
class	Polypodiopsida

Rank	Name	Author
family	Marsileaceae	Mirbel 1802
family	Marsileaceae	Mirbel 1802

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Marsileaceae Mirbel 1802

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G. †Collarisporites Déak 1964

†Collarisporites fuscus Déak 1964

G. †Crybelosporites Dettmann 1963

†Crybelosporites australis Archangelsky and Llorens 2005

†Crybelosporites bellus Singh 1983

†Crybelosporites berberioides Burger 1976

†Crybelosporites brenneri Playford 1971

†Crybelosporites mamelonatus Lima 1978

†Crybelosporites pannuceus Srivastava 1977

†Crybelosporites punctatus Dettmann 1963

†Crybelosporites striatus Dettmann 1963

†Crybelosporites stylosus Dettmann 1963

†Crybelosporites vectensis Kemp 1970

G. †Gabonisporis Boltenhagen 1967

†Gabonisporis cristata Sweet 1986

†Gabonisporis vigourouxii Boltenhagen 1967

G. †Lugiomarsiglia Hermsen et al. 2013

†Lugiomarsiglia aquatica Hermsen et al. 2013

G. Marsilea Linnaeus 1753 [water clover]

†Marsilea andersoni Hollick 1904

†Marsilea attenuata Lesquereux 1875

†Marsilea intertrappea Patil and Upadhaye 1979

†Marsilea johnhallii Skog and Dilcher 1992

G. †Marsileaceaephyllum Nagalingum 2007

†Marsileaceaephyllum johnhallii Skog and Dilcher 1992

G. †Mirasolita Hermsen et al. 2013

†Mirasolita irupensis Hermsen et al. 2013

G. †Molaspora Hall 1963

†Molaspora lobata Hall and Peake 1968

G. †Pillularia Linneo

†Pillularia novaezealandiae Couper 1953

G. Regnellidium Lindman

†Regnellidium thomastaylorii Cúneo et al. 2013

G. †Rhizocarpites Heer 1878

†Rhizocarpites singularis Heer 1878

G. †Rodeites Sahni 1943

†Rodeites dakshini Sahni 1943

†Rodeites intertrappeana Chitaley and Paradkar 1972

†Rodeites polycarpa Chitaley and Paradkar 1973

Invalid names: Cystorrhiza Massalongo 1859 [nomen nudum]

Diagnosis

Reference	Diagnosis
E. J. Hermsen et al. 2013	Diagnosis of marsileaceous vegetative organs. Rhizomes bearing roots and simple, unbranched filiform or compound leaves, or isolated compound leaves. Compound leaves petiolate with two or four leaflets (sometimes more) borne oppositely or in a pseudowhorl at the petiole apex. Leaflet venation of a single order, dichotomous with anastomoses, midvein absent; frequency of anastomoses variable (venation ranging from nearly lacking anastomoses to being fully reticulate). Areoles elongate lengthwise (but may become shorter distally) and generally fusiform in shape, lacking freely-ending veinlets.