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Grallator ssatoi

Osteichthyes - Grallatoridae

Taxonomy
Jeholosauripus ssatoi was named by Yabe et al. (1940) [originally spelled "s-satoi"]. It is considered to be a form taxon. Its type specimen is TIU 61677, a footprint, and it is a trace fossil. Its type locality is Ssuchiatzu tracksite, which is in a Kimmeridgian/Valanginian terrestrial sandstone/conglomerate in the Tuchengzi Formation of China.

It was recombined as Grallator ssatoi by Zhen et al. (1989), Lockley and Matsukawa (1998), Peng et al. (2005), Fujita et al. (2008), Xing et al. (2009), Sullivan et al. (2009), Xing et al. (2011), Lockley et al. (2013), Lockley et al. (2014), Xing et al. (2015), Xing et al. (2015), Li et al. (2022).

Sister species lacking formal opinion data

Synonymy list
YearName and author
1940Jeholosauripus ssatoi Yabe et al. p. 169–170 fig. 1
1941Jeholosauripus ssatoi Noda and Shikama p. 477
1942Jeholosauripus ssatoi Shikama pp. 23-25
1958Jeholosauripus ssatoi Kuhn p. 27
1960Jeholosauripus ssatoi Young p. 54
1963Jeholosauripus ssatoi Kuhn p. 100
1971Jeholosauripus ssatoi Haubold p. 68
1974Jeholosauripus ssatoi Haubold p. 138
1984Jeholosauripus ssatoi Cheng p. 222
1989Grallator ssatoi Zhen et al. p. 188
1991Jeholosauripus ssatoi Azuma and Takeyama p. 48
1998Grallator ssatoi Lockley and Matsukawa p. 136
2005Grallator ssatoi Peng et al.
2008Grallator ssatoi Fujita et al. p. 41 fig. 10
2009Grallator ssatoi Sullivan et al. p. 36
2009Jeholosauripus ssatoi Xing et al. p. 23
2009Grallator ssatoi Xing et al. p. 708
2011Grallator ssatoi Xing et al. p. 431
2013Grallator ssatoi Lockley et al. p. 10
2014Grallator ssatoi Lockley et al. p. 775
2015Grallator ssatoi Xing et al. p. 1011
2015Grallator ssatoi Xing et al. p. 17
2022Grallator ssatoi Li et al. p. 4232

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
RankNameAuthor
subclassEureptilia()
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Theropoda()
familyGrallatoridae
genusGrallator
speciesssatoi()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Grallator ssatoi Yabe et al. 1940
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Diagnosis
ReferenceDiagnosis
S. Zhen et al. 1989Plantigrade, tridactyl, but no trace of a hallux, lateral digits, nor caudal impression. General outline deltoid. Proximal end of III starting much anterior to that of II and IV. Size rather small: length of footprints is 7.0–12.0 cm and the width is 5.0–8.5 cm. The angles between the digits are II-14°-III-13°-IV.
G. Peng et al. 2005Three-toed footprints made by small, bipedal and digitigrate coelurosaurs. Length of each footprint is about twice of its width. The divarication between digit II and III is 14°~29°, and that between digit III and IV is 13°~27°. Digit III is much longer than digit II and IV. No impressions of hallux and tail.