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Nuculanidae (pointed nut clam)
Taxonomy
Nuculanidae was named by Adams and Adams (1858) [m. kosnik taxon no: 5054; =Ledidae H. Adams and A. Adams 1858; Poroledidae Scarlato and Starobogatov 1979; Ledidae published in Jan. 1858, Nuculanidae published in Nov. 1858, Nuculanidae maintained under ICZN Code Art. 40b.]. It is extant. Its type is Nuculana.
It was assigned to Pelecypoda by Turner (1938); to Nuculacea by Adams and Adams (1858), Grant, IV and Gale (1931), Gardner (1945), DuBar (1958); to Nuculanacea by Newell (1965), Vokes (1980), Dockery (1982), Squires (1984), Ward and Blackwelder (1987), Squires (1987), Stilwell and Zinsmeister (1992), Maxwell (1992); to Nuculoida by Givens and Kennedy (1976), Squires (1999), Spencer et al. (2004), El Qot (2006); and to Nuculanoidea by Coan et al. (2000), Todd (2001), Stilwell et al. (2004), Fang and Cope (2004), Bieler and Mikkelsen (2006), Fang (2006), Mikkelsen and Bieler (2008), Nevesskaja (2009), Bouchet et al. (2010), Carter et al. (2011), Coan and Valentich-Scott (2012), Ros-Franch et al. (2014).
It was assigned to Pelecypoda by Turner (1938); to Nuculacea by Adams and Adams (1858), Grant, IV and Gale (1931), Gardner (1945), DuBar (1958); to Nuculanacea by Newell (1965), Vokes (1980), Dockery (1982), Squires (1984), Ward and Blackwelder (1987), Squires (1987), Stilwell and Zinsmeister (1992), Maxwell (1992); to Nuculoida by Givens and Kennedy (1976), Squires (1999), Spencer et al. (2004), El Qot (2006); and to Nuculanoidea by Coan et al. (2000), Todd (2001), Stilwell et al. (2004), Fang and Cope (2004), Bieler and Mikkelsen (2006), Fang (2006), Mikkelsen and Bieler (2008), Nevesskaja (2009), Bouchet et al. (2010), Carter et al. (2011), Coan and Valentich-Scott (2012), Ros-Franch et al. (2014).
Subtaxa
Acilana, Acutispinula, Anthracoleda, Australoportlandia, Bathyspinula, Calorhadia, Clidophorus, Costatoleda, Costelloleda, Dacromya, Ezonuculana, Indonuculana (syn. Reticulana), Ledella (syn. Junonia), Ledellinae, Litorhadia, Mesosaccella, Multidentata, Nuculana (syn. Crassoleda, Leda) (type genus), Nuculana gabbii, Nuculaninae (syn. Adraninae), Orthoyoldia, Phestioidea, Poroleda (syn. Lamellileda), Poroledidae, Praeportlandia, Propeleda, Saccella, Scissuladrana, Spineilo, Veteranellinae
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1858 | Ledidae Adams and Adams |
| 1858 | Nuculanidae Adams and Adams |
| 1860 | Leda protexta Gabb p. 393 figs. pl., 48, fig. 23 |
| 1864 | Leda protexta Gabb p. 199 figs. PL 20, Fig. 185 |
| 1866 | Nuculana gabbii Conrad p. 3 |
| 1869 | Leda gabbi Gabb pp. 197 - 198 |
| 1896 | Leda gabbi Stanton p. 1041 |
| 1904 | Ledidae Glenn p. 395 |
| 1915 | Leda vaderensis Dickerson p. 52 figs. pl. 1, fig. 3 |
| 1916 | Ledidae Raymond |
| 1917 | Leda gabbi Waring p. 76 figs. Pl. 13, Fig. 6 |
| 1923 | Ledidae Clark and Arnold p. 137 |
| 1925 | Leda vogdesi Anderson and Hanna pp. 177 - 179 figs. pl. 2, figs. 8-9 |
| 1925 | Leda gabbi Nelson p. 404 |
| 1927 | Leda gabbi Clark and Woodford pp. 85 - 86 figs. Plate 14, figure 2 |
| 1927 | Leda vogdesi Hanna p. 271 |
| 1930 | Saccella gabbii Stewart pp. 55 - 56 figs. Plate fig. and pl. 10, fig. 4 |
| 1931 | Nuculanidae Grant, IV and Gale p. 116 |
| 1931 | Ledidae Olsson p. 36 |
| 1938 | Nuculana gabbi Turner p. 13 |
| 1938 | Nuculanidae Turner p. 41 |
| 1945 | Nuculanidae Gardner |
| 1958 | Nuculanidae DuBar p. 153 |
| 1965 | Nuculanidae Newell p. 16 |
| 1976 | Nuculanidae Givens and Kennedy p. 973 |
| 1980 | Nuculanidae Vokes p. 5 |
| 1982 | Ledellinae Allen and Sanders |
| 1982 | Nuculanidae Dockery p. 28 |
| 1983 | Veteranellinae Chen et al. |
| 1983 | Nuculana (Saccella) gabbii Moore p. A16 figs. Plate 2, figures 7, 8 |
| 1984 | Nuculana (Saccella) gabbii Squires p. 41 figs. Figure 10b |
| 1984 | Nuculanidae Squires p. 41 |
| 1987 | Nuculanidae Squires p. 54 |
| 1987 | Nuculanidae Ward and Blackwelder p. 129 |
| 1992 | Nuculanidae Maxwell p. 56 |
| 1992 | Nuculanidae Stilwell and Zinsmeister p. 48 |
| 1999 | Nuculana (Saccella) gabbii Squires p. 23 figs. Figure 50 |
| 1999 | Nuculanidae Squires p. 23 |
| 2000 | Ledellinae Coan et al. pp. 82, 94 |
| 2000 | Nuculanidae Coan et al. pp. 82, 83 |
| 2001 | Nuculanidae Todd |
| 2004 | Nuculanidae Fang and Cope |
| 2004 | Nuculanidae Spencer et al. |
| 2004 | Nuculanidae Stilwell et al. p. 17 |
| 2006 | Nuculanidae Bieler and Mikkelsen p. 228 |
| 2006 | Nuculanidae El Qot p. 17 |
| 2006 | Nuculanidae Fang |
| 2008 | Nuculanidae Mikkelsen and Bieler p. 19 |
| 2009 | Nuculanidae Nevesskaja |
| 2010 | Nuculanidae Bouchet et al. |
| 2011 | Nuculanidae Carter et al. p. 6 |
| 2012 | Nuculanidae Coan and Valentich-Scott p. 57 |
| 2014 | Nuculanidae Ros-Franch et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Nuculanidae Adams and Adams 1858 [pointed nut clam]
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G. †Acilana Khomenko 1937
G. †Anthracoleda Benedetto 1980
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Subg. †Anthracoleda (Pseudoleda) Sanchez et al. 1995
+
†Anthracoleda (Pseudoleda) minuta Sanchez et al. 1995
G. Bathyspinula Filatova 1958
+
Subg. Bathyspinula (Bathyspinula) Allen and Sanders 1982
Spinula (Bathyspinula) oceanica Filatova 1958
G. Costatoleda
G. Costelloleda Hertlein and Strong 1940
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†Nuculana (Costelloleda) powersi Dickerson 1914
†Nuculana (Costelloleda) whitensis Farinati 1978
G. †Dacromya Agassiz 1840
+
†Dacromya chetaensis Sanin 1976
†Dacromya nagaensis Awad 1945
†Dacryomya elongata Fan 1963
†Dacryomya kiparisovae Kurushin 1990
†Dacryomya peculiaris Kurushin 1990
†Dacryomya polaris Kiparisova 1937
†Dacryomya skorochodi Kiparisova 1938
†Dacryomya tarua Skwarko 1973
G. †Indonuculana Kanjilal and Singh 1980
+
Invalid names: Reticulana Kanjilal and Singh 1973 [synonym]
G. Ledella Verrill and Bush 1897
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Subg. Ledella (Amphilata) Filatova and Schileyko 1984
+
Ledella (Amphilata) kermadecensis Knudson 1970
Ledella (Amphilata) librata Dell 1952
Subg. Ledella (Ledella) Verrill and Bush 1897
Ledella herdmani Dell 1953
†Ledella leptorhyncha Tate 1886
Ledella praelonga Tate 1886
Ledella sublevis Verrill and Bush 1898 [Polished Nut Clam]
Invalid names: Junonia Seguenza 1877 [synonym]
G. †Litorhadia Stewart 1930
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Calorhadia (Litorhadia) acala Dall 1898
†Calorhadia (Litorhadia) bastropensis Harris 1895
†Calorhadia (Litorhadia) compsa Gabb 1860
†Calorhadia (Litorhadia) evanescentior Stenzel and E. K. 1957
†Calorhadia (Litorhadia) lisbonensis Aldrich 1895
†Calorhadia (Litorhadia) mater Meyer 1885
†Calorhadia (Litorhadia) santaanai Gardner 1945
†Litorhadia astoriana Henderson 1920
+
Invalid names: Leda subimpressa Howe 1922 [synonym], Nucula impressa Conrad 1849 [replaced]
†Litorhadia buenoi Etayo-Serna 1979
G. †Mesosaccella Chavan 1946
+
†Mesosaccella partimstriata Wood 1864
†Mesosaccella perdita Conrad 1852
†Mesosaccella rogovi Hryniewicz et al. 2014
†Mesosaccella toddi Hryniewicz et al. 2014
G. †Multidentata Krishtofovich 1964
G. Nuculana Link 1807
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Subg. †Nuculana (Borissia)
Subg. Nuculana (Costanuculana) Habe 1951
Subg. †Nuculana (Dacryomya) Agassiz 1840
Subg. Nuculana (Nuculana) Link 1807
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†Nuculana (Nuculana) arenicola Kendrick and Vartak 2007
Nuculana (Nuculana) confusa Hanley 1860
Nuculana (Nuculana) elongorostrata Clark 1918
Nuculana (Nuculana) fossa Baird 1863
Nuculana (Nuculana) fossa sculpta Dall 1916
Nuculana (Nuculana) hamata Carpenter 1864
Nuculana (Nuculana) leonina Dall 1896
+
Invalid names: Leda amiata Dall 1916 [synonym], Leda liogona Dall 1916 [synonym]
Nuculana (Nuculana) minuta Müller 1776
†Nuculana (Nuculana) naibandensis Hautmann 2001
Nuculana (Nuculana) navisa Dall 1916
†Nuculana (Nuculana) odiyamensis Kendrick and Vartak 2007
†Nuculana (Nuculana) senni Kugler et al. 1984
Nuculana (Nuculana) tenuisulcata Couthouy 1838
Invalid names: Nuculana (Nuculana) minuta praecursor Arnold 1903 [synonym]
Subg. Nuculana (Politoleda) Hertlein and Strong 1940
+
†Nuculana (Politoleda) forcarti Jung 1965
Politoleda polita Sowerby 1833
Subg. Nuculana (Scaeoleda) Iredale 1924
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Nuculana (Scaeoleda) verconis Tate 1891
†Scaeoleda acinaciformis Tate 1886
†Scaeoleda crebricostata Tenison Woods 1877
†Scaeoleda woodsii Tate 1886
Subg. Nuculana (Tenuileda) Habe 1977
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Subg. Nuculana (Thestyleda) Iredale 1929
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Nuculana (Thestyleda) ramsayi Smith 1885
Thestyleda investigator Dell 1952
†Nuculana acutilineata Hybertsen and Kiel 2018
†Nuculana akidota Dockery 1982
†Nuculana alpina Winkler 1859
†Nuculana andrewi Marwick 1931
†Nuculana antarctirostrata Stilwell et al. 2004
†Nuculana asiatica Kurushin 1985
†Nuculana biangulata Easton 1962
Nuculana concentrica Say 1824 [Concentric Nut Clam]
†Nuculana cowlitzensis Weaver and Van Winkle Palmer 1922
†Nuculana cultellus von Koenen 1897
Nuculana cuneata Sowerby 1833
†Nuculana curvirostris de Cserna 1961
Nuculana decora Adams 1856
†Nuculana dewalquei Terquem and Piette 1868
†Nuculana doris d'Orbigny 1849
†Nuculana elmana Etherington 1931
Nuculana emarginata Lamarck 1819
†Nuculana etheridgei Maxwell 1964
†Nuculana excavata Goldfuss 1837
†Nuculana fluvilutea Marwick 1931
†Nuculana forcarti Jung 1965
†Nuculana gnomon Jung 1965
†Nuculana gracillima Jung 1965
†Nuculana grasslei Allen 1993
Nuculana jamaicensis d'Orbigny 1853 [Jamaican Nut Clam]
†Nuculana konishii Hayami 1961
†Nuculana liciata Conrad 1843
†Nuculana longifrons Conrad 1860
†Nuculana mariae d'Orbigny 1844
†Nuculana miaocunensis Chen 1976
†Nuculana minima Sowerby 1812
Nuculana minuta Müller 1776
†Nuculana moriyai Marincovich, Jr. 1993
Nuculana oxia Dall 1916
†Nuculana oxyrhyncha Philippi 1887
†Nuculana parkei Anderson and Hanna 1925
†Nuculana parva Jung 1965
Nuculana penita Conrad 1848
Nuculana pernula Müller 1779
+
Invalid names: Leda buccata Müller 1842 [synonym], Nuculana jacksoni Gould 1841 [synonym], Nuculana rostrata Bruguiére 1789 [synonym]
†Nuculana posterolaevia Kiel and Goedert 2007
†Nuculana probellula Marwick 1929
†Nuculana redondoensis Burch 1944
†Nuculana rostellata Conrad 1841
†Nuculana rostratruncata Gardner 1916
Nuculana semen Smith 1885 [Seed Nut Clam]
†Nuculana semicrenulata Trechmann 1917
†Nuculana socialis Stoliczka 1871
†Nuculana thompsoni Cox 1965
†Nuculana timorensis Krumbeck 1924
†Nuculana triangulata Meyer 1886
†Nuculana undulostriata Fletcher 1945
Nuculana verrilliana Dall 1886 [Barbed Nut Clam]
†Nuculana vexillata Marwick 1953
†Nuculana virgo Martin 1879
Nuculana vitrea d'Orbigny 1853 [Glassy Nut Clam]
†Nuculana waterhousei Etheridge 1888
†Nuculana waynensis Mansfield 1940
†Nuculana whitmani Dall 1909
Nuculana willamettensis Shumard 1858
†Nuculana willetti Marwick 1953
†Nuculana zieteni Brauns 1871
Invalid names: Crassoleda Savitskii 1969 [synonym], Leda Schumacher 1817 [synonym]
Subfm. Nuculaninae Adams and Adams 1858
+
G. Adrana Adams and Adams 1858
+
†Adrana aegra Olsson 1964
†Adrana agronomica Maury 1924
†Adrana aldrichiana Harris 1895
†Adrana dalliana Olsson 1922
†Adrana electa Adams 1858
†Adrana ensinoides Olsson 1922
Adrana exoptata Pilsbry and Lowe 1932
Adrana lanceolata Sowerby 1817
†Adrana quitanensis Olsson 1922
Adrana sowerbyana d'Orbigny 1833
†Adrana stena Woodring 1973
G. †Culunana Lintz 1958
G. Jupiteria Bellardi 1875
+
Subg. †Jupiteria (Surojupiteria) Stilwell and Zinsmeister 1992
+
†Jupiteria (Surojupiteria) dissensa Stilwell and Zinsmeister 1992
†Jupiteria aipeia Woodring 1973
Jupiteria cellulita Dall 1896
Jupiteria concava Bronn 1831
†Jupiteria solida Dall 1886
†Jupiteria subtumida Woodring 1925
†Leda (Jupiteria) bowdenensis Woodring 1925
†Leda (Jupiteria) bowdenensis bowdenensis Woodring 1925
†Leda (Jupiteria) duerdeni Woodring 1925
†Nuculana (Jupiteria) asiatica Kurushin 1985
Nuculana (Jupiteria) delliana Huber 2010
†Nuculana (Jupiteria) kapua Marwick 1931
†Nuculana (Jupiteria) karaka Marwick 1931
†Nuculana (Jupiteria) kouhaiensis Marwick 1931
†Nuculana (Jupiteria) leachi Marwick 1931
†Nuculana (Jupiteria) nugax Marwick 1931
Nuculana (Jupiteria) penderi Dall and Bartsch 1910
†Nuculana (Jupiteria) prisca Deshayes 1858
†Nuculana (Jupiteria) totangiensis Marwick 1931
†Nuculana (Jupiteria) uawa Marwick 1931
Invalid names: Epacroleda Savitskii 1969 [synonym]
G. †Ledina Dall 1898
+
†Ledina duttonae Vokes 1939
†Ledina fresnoensis Dickerson 1916
†Ledina smirna Dall 1898
G. †Pseudoportlandia Woodring 1925
+
Pseudoportlandia clara Guppy 1873
†Pseudoportlandia tahuia Marwick 1942
Invalid names: Adraninae Scarlato and Starobogatov 1979 [synonym]
G. Orthoyoldia Verrill 1897
+
†Orthoyoldia claibornensis Conrad 1848
Orthoyoldia crosbyana Guppy 1882
†Orthoyoldia psammotaea Dall 1898
†Yoldia (Orthoyoldia) haboroensis Uozumi 1957
†Yoldia (Orthoyoldia) quiba Olsson 1942
G. Poroleda Hutton 1893
+
Poroleda lanceolata Hutton 1885
Invalid names: Lamellileda Cotton 1930 [synonym]
G. †Praeportlandia Savitskii 1969
G. Propeleda Iredale 1924
+
†Nuculana (Propeleda) darwini Philippi 1887
†Nuculana (Propeleda) dorbignyi Philippi 1887
†Nuculana (Propeleda) medinae Philippi 1887
Propeleda carpenteri Dall 1881 [Carpenter's Smooth Nut Clam]
†Propeleda trulliformis Marwick 1931
G. Saccella Woodring 1925
+
†Nuculana (Saccella) alaeformis Gabb 1869
†Nuculana (Saccella) andrewi Marwick 1931
†Nuculana (Saccella) arowhana Marwick 1931
†Nuculana (Saccella) bolivarensis Clark and Durham 1946
†Nuculana (Saccella) cahillensis Arnold 1908
†Nuculana (Saccella) chaneyi Vokes 1939
†Nuculana (Saccella) chehalisensis Weaver 1912
†Nuculana (Saccella) costulata Deshayes 1858
†Nuculana (Saccella) crassatelloides Olsson 1964
†Nuculana (Saccella) fabata Nelson 1925
†Nuculana (Saccella) furlongi Trask 1922
†Nuculana (Saccella) gabbii Conrad 1866
+
Invalid names: Leda protexta Gabb 1860 [synonym], Leda vaderensis Dickerson 1915 [synonym], Leda vogdesi Anderson and Hanna 1925 [synonym]
†Nuculana (Saccella) galeottiana Nyst 1843
†Nuculana (Saccella) gnomon Jung 1965
†Nuculana (Saccella) gracillima Jung 1965
†Nuculana (Saccella) hamata Eames 1951
†Nuculana (Saccella) hondana Vokes 1939
†Nuculana (Saccella) karlmartini Weisbord 1964
†Nuculana (Saccella) markleyensis Clark 1918
†Nuculana (Saccella) merriami Dickerson 1917
†Nuculana (Saccella) nolfi Glibert 1985
†Nuculana (Saccella) ochsneri Anderson and Martin 1914
†Nuculana (Saccella) packardi Dickerson 1914
†Nuculana (Saccella) pakistanica Eames 1951
†Nuculana (Saccella) parva Jung 1965
†Nuculana (Saccella) pseudoscissurata Takeda 1953
†Nuculana (Saccella) pulchrisinuosa Clark 1918
†Nuculana (Saccella) punjabensis Eames 1951
†Nuculana (Saccella) rakhiensis Eames 1951
†Nuculana (Saccella) ramonensis Clark 1918
†Nuculana (Saccella) semibistriata Eames 1951
†Nuculana (Saccella) silvestris Cotter 1923
†Nuculana (Saccella) soriensis Eames 1951
†Nuculana (Saccella) striata Lamarck 1805
†Nuculana (Saccella) taylorensis Mansfield 1940
†Nuculana (Saccella) umboniinflata Eames 1951
†Nuculana (Saccella) waikohuensis Marwick 1931
†Nuculana (Saccella) washingtonensis Weaver 1916
+
Invalid names: Leda lincolnensis Weaver 1916 [synonym]
†Nuculana ochsneri elmana Etherington 1931
†Saccella acrita Dall 1908
†Saccella acrita epacra Woodring 1973
†Saccella acuminata von Buch 1845
†Saccella acuticauda Pritchard 1901
†Saccella amelga Moore 1963
†Saccella balboae Brown and Pilsbry 1911
†Saccella calkinsi Moore 1963
†Saccella canonica Dall 1890
†Saccella charanica Olsson 1931
†Saccella chiriquiana Olsson 1942
†Saccella davidana Olsson 1942
†Saccella davilae Olsson 1922
Saccella eburnea Sowerby 1833
Saccella elenensis Sowerby I 1833
+
Invalid names: Leda acapulcensis Pilsbry and Lowe 1932 [synonym]
†Saccella fabalis Olsson 1964
Saccella fastigata Keen 1958
†Saccella fontinalis Pritchard 1901
Saccella hindsii Hanley 1860
†Saccella hokkaidoensis Oyama and Mizuno 1958
†Saccella incognita Guppy 1867
Saccella laeviradius Pilsbry and Lowe 1932
Saccella maxwelli Beu 2006
†Saccella miradorana Olsson 1931
†Saccella orcutti Arnold 1908
Saccella ornata d'Orbigny 1845
†Saccella packardi Forbes 1848
†Saccella parva Rogers and Rogers 1837
†Saccella peltella Dall 1903
†Saccella peruviana Dall 1895
†Saccella phlyctaena Woodring 1973
†Saccella quercollis Harris 1896
†Saccella semiteres Hutton 1877
†Saccella subcerata Woodring 1925
†Saccella tenellula Bartrum and Powell 1928
†Saccella uvasana Dickerson 1915
†Saccella vulgaris Brown and Pilsbry 1913
+
Invalid names: Leda obliterata Brown and Pilsbry 1913 [synonym]
†Saccella westendorpi Nyst and Westendorp 1839
+
Invalid names: Nuculana gracilis Deshayes 1860 [synonym]
Sacella atakta Gardner 1927
G. Scissuladrana Petuch 1987
G. †Spineilo Finlay and Marwick 1937
Invalid names: Ledellinae Allen and Sanders 1982 [invalid subgroup], Ledidae Adams and Adams 1858 [synonym], Nuculana gabbii Conrad 1866 [nomen nudum], Poroledidae Scarlato and Starobogatov 1979 [empty], Veteranellinae Chen et al. 1983 [invalid subgroup]
Diagnosis
No diagnoses are available