PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Dyrosaurus phosphaticus

Reptilia

Taxonomy

Crocodilus phosphaticus was named by Thomas (1893). Its type specimen is MNHN 1901-11, a partial skeleton (one tooth, one caudal vertebra, one distal extremity of an ulna and one distal extremity of a radius), and it is a 3D body fossil. Its type locality is Djebel Teldja, which is in a Ypresian marine phosphorite in Tunisia.

It was recombined as Dyrosaurus phosphaticus by Stefano (1903), Buffetaut (1978), Moody and Buffetaut (1981), Jouve (2005), Jouve et al. (2006), Bronzati et al. (2012) and Sertich and O'Connor (2014).

Sister species lacking formal opinion data

D. choumovitchi

Synonyms

Dryosaurus thevestensis was named by Pomel (1894).

It was synonymized subjectively with Dryosaurus phosphaticus by Jouve (2005).
Dyrosaurus choumowitchi was named by Bergounioux (1955).

It was synonymized subjectively with Dyrosaurus phosphaticus by Buffetaut (1978); it was synonymized subjectively with Dryosaurus phosphaticus by Jouve (2005).
Dryosaurus berneti was named by Bergounioux (1955).

It was synonymized subjectively with Dryosaurus phosphaticus by Jouve (2005).
Dryosaurus variabilis was named by Bergounioux (1955).

It was synonymized subjectively with Dryosaurus phosphaticus by Jouve (2005).
Dyrosaurus berneti was named by Bergounioux (1955). It is a 3D body fossil. Its type locality is Metlaoui, which is in a Ypresian coastal phosphorite in the Gafsa Phosphates Formation of Tunisia.

It was synonymized subjectively with Dyrosaurus phosphaticus by Buffetaut (1978) and Jouve (2005).
Dyrosaurus variabilis was named by Bergounioux (1955). It is a 3D body fossil. Its type locality is Metlaoui, which is in a Ypresian coastal phosphorite in the Gafsa Phosphates Formation of Tunisia.

It was considered a nomen dubium by Buffetaut (1978); it was synonymized subjectively with Dyrosaurus phosphaticus by Jouve (2005).

Synonymy list

Year	Name and author
1894	Dryosaurus thevestensis Pomel p. 1310
1903	Dyrosaurus phosphaticus Stefano p. 54
1955	Dryosaurus berneti Bergounioux p. 1917
1955	Dyrosaurus berneti Bergounioux p. 1917
1955	Dyrosaurus choumowitchi Bergounioux p. 1917
1955	Dyrosaurus variabilis Bergounioux p. 1917
1955	Dryosaurus variabilis Bergounioux p. 1918
1978	Dyrosaurus phosphaticus Buffetaut p. 238
1981	Dyrosaurus phosphaticus Moody and Buffetaut p. 127
2005	Dyrosaurus phosphaticus Jouve p. 324
2006	Dyrosaurus phosphaticus Jouve et al.
2012	Dyrosaurus phosphaticus Bronzati et al.
2014	Dyrosaurus phosphaticus Sertich and O'Connor

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()

Rank	Name	Author
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
—	Pseudosuchia	(Zittel 1890)
—	Suchia	Krebs 1974
—	Paracrocodylomorpha
—	Loricata	(Merrem 1820)
—	Crocodylomorpha	()
—	Solidocrania
suborder	Crocodyliformes
—	Mesoeucrocodylia	Whetstone and Whybrow 1983
—	Neosuchia	Clark 1988
—	Coelognathosuchia	()
—	Tethysuchia	()
—	Dyrosauroidea
—	Dyrosauridae	()
—	Hyposaurinae	()
genus	Dyrosaurus	Pomel 1894
species	phosphaticus	(Thomas 1893)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Dyrosaurus phosphaticus Thomas 1893

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Invalid names: Dyrosaurus berneti Bergounioux 1955 [synonym], Dyrosaurus variabilis Bergounioux 1955 [synonym]

Diagnosis

Reference	Diagnosis
S. Jouve 2005	Long and slender snout, about 68% of the total skull length; posterior wall of the supratemporal fenestra almost vertical and poorly exposed in dorsal view; inter- fenestral bar moderately wide and slightly T-shaped in trans- verse section; posterior margin of the skull roof straight in dorsal view; occipital tuberosities moderately extended; small posterolateral wing of the palatine within the suborbital fenestra; most anterior point of the posterior margin of the pterygoid wings far anterior to the level of the medial eusta- chian foramen, at mid-length between the foramen and the posterior margin of the choanal septum; deep craniocaudally elongated sulcus between the posterior margin of the choanae and the medial eustachian foramen.

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

marineuc

Locomotion:

actively mobilec

Diet:

carnivoresubo

Created:

2017-04-17 10:39:01

Modified:

2017-04-17 10:39:01

Source: subo = suborder, c = class, subp = subphylum, uc = unranked clade

References: Benton 1983, Carroll 1988, Hendy et al. 2009, Bush and Bambach 2015

Age range: Early/Lower Eocene or 55.80000 to 48.60000 Ma

Collections (4 total)

Time interval	Ma	Country or state	Original ID and collection number
Ypresian	56.0 - 47.8	Tunisia	Crocodilus phosphaticus (type locality: 136905) Dyrosaurus phosphaticus, Dyrosaurus berneti, Dyrosaurus variabilis (106501)
Ypresian	56.0 - 47.8	Algeria	Dyrosaurus phosphaticus (136906)
Early/Lower Eocene	55.8 - 48.6	Tunisia (Gafsa)	Dyrosaurus phosphaticus (151620)