PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Coccolithophyceae

Taxonomy

Coccolithophyceae was named by Rothmaler (1951). It is extant.

It was assigned to Haptophyta by Silva et al. (2007).

Sister taxa

Calyptrosphaeraceae, Ceratolithaceae, Ellipsagelosphaeraceae, Goniolithaceae, Lapideacassaceae, Rhomboasteraceae

Synonyms

Prymnesiophyceae was named by Hibberd (1976). It is extant.

It was synonymized subjectively with Coccolithophyceae by Silva et al. (2007).

It was assigned to Haptophyta by Young and Bown (1997).

Synonymy list

Year	Name and author
1951	Coccolithophyceae Rothmaler
1976	Prymnesiophyceae Hibberd
1997	Prymnesiophyceae Young and Bown
2007	Coccolithophyceae Silva et al. p. 473

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Rank	Name	Author
—	Life
—	Eukaryota	()
—	Chromalveolata

Rank	Name	Author
phylum	Haptophyta	Hibberd ex Cavalier-Smith 1986
class	Coccolithophyceae
class	Coccolithophyceae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Cl. Coccolithophyceae Rothmaler 1951

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G. †Athenagalea Hattner and Wise 1980

G. †Biantholithus Bramlette and Martini 1964

Or. Braarudosphaerales Aubry 2013

Fm. †Polycyclolithaceae Forchheimer 1972

G. †Assipetra Roth 1973

†Assipetra infracretacea Thierstein 1973

†Assipetra terebrodentarius Applegate and Bralower 1987

†Assipetra terebrodentarius youngii Tremolada and Erba 2002

†Assipetra terebrodentarius terebrodentarius Applegate and Bralower 1987

G. †Eprolithus Stover 1966

†Eprolithus antiquus Perch-Nielsen 1979

†Eprolithus apertior Black 1973

†Eprolithus floralis Stradner 1962

†Eprolithus moratus Stover 1966

†Eprolithus octopetalus Varol 1992

†Eprolithus rarus Varol 1992

Invalid names: Polycyclolilthus Forchheimer 1968 [synonym]

G. †Farhania Varol 1992

†Farhania varolii Jakubowski 1986

G. †Hayesites Manivit 1971

†Hayesites albiensis Manivit 1971

†Hayesites irregularis Thierstein et al. 1972

G. †Hexalithus Gardet 1955

†Hexalithus gardetiae Bukry 1969

†Hexalithus noeliae Loeblich and Tappan 1966

G. †Lithastrinus Stradner 1962

†Lithastrinus grillii Stradner 1962

†Lithastrinus pentabrachius Varol 1992

†Lithastrinus quadricuspis Farhan 1997

†Lithastrinus septenarius Forchheimer 1972

G. †Micula Vekshina 1959 [coccolithophore]

†Micula adumbrata Burnett 1998

†Micula clypeata Lees and Bown 2005

†Micula concava Stradner 1960

†Micula cubiformis Forchheimer 1972

†Micula decussata Vekshina 1959 [coccolithophore]

†Micula murus Martini 1961

†Micula praemurus Bukry 1973

†Micula premolisilvae Lees and Bown 2005

†Micula prinsii Perch-Nielsen 1979

†Micula staurophora Gardet 1955

†Micula swastica Stradner and Steinmetz 1984

Invalid names: Nannotetraster Martini and Stradner 1960 [synonym]

G. †Polycostella Thierstein 1971

†Polycostella beckmanii Thierstein 1971

†Polycostella senaria Thierstein 1971

G. †Quadrum Prins and Perch-Nielsen 1977

†Quadrum bengalensis Burnett 1998

†Quadrum eneabrachium Varol 1992

†Quadrum eptabrachium Varol 1992

†Quadrum gartneri Prins and Perch-Nielsen 1977

†Quadrum giganteum Varol 1992

†Quadrum intermedium Varol 1992

†Quadrum octobrachium Varol 1992

†Quadrum svabenickae Burnett 1998

G. †Radiolithus Stover 1966

†Radiolithus hollandicus Varol 1992

†Radiolithus orbiculatus Forchheimer 1972

†Radiolithus planus Stover 1966

G. †Rucinolithus Stover 1966

†Rucinolithus magnus Bukry 1975

†Rucinolithus windleyae Rutledge and Bown 1996

†Rucinolithus wisei Thierstein 1971

G. †Tegulalithus Crux 1986

†Tegulalithus septentrionalis Stradner 1963

†Tegulalithus tessellatus Stradner 1968

G. †Uniplanarius Hattner and Wise 1980

†Uniplanarius clarkei Lees and Bown 2005

†Uniplanarius gothicus Deflandre 1959

†Uniplanarius sissinghii Perch-Nielsen 1986

†Uniplanarius trifidus Stradner 1961

Fm. Ceratolithaceae Norris 1965

G. Amaurolithus Gartner and Bukry 1975

Amaurolithus primus Bukry and Percival 1971

G. Angulolithina Bukry 1973

G. Ceratolithus Kamptner 1950

G. Florisphaera Okada and Honjo 1973

G. †Nicklithus Raffi et al. 1998

Nicklithus amplificus Bukry and Percival 1971

G. †Orthorhabdus Bramlette and Wilcoxon 1967

Or. †Coccolithales Schwarz 1932

Fm. Coccolithaceae Poche 1913

G. †Bicolumnus Wei and Wise 1990

G. †Birkelundia Perch-Nielsen 1971

G. †Bramletteius Gartner 1969

G. †Campylosphaera Kamptner 1963

G. †Chiasmolithus Hay et al. 1966

†Chiasmolithus bidens Bramlette and Sullivan 1961

†Chiasmolithus consuetus Bramlette and Sullivan 1961

†Chiasmolithus danicus Hay and Mohler 1967

†Chiasmolithus eograndis Perch-Nielsen 1971

†Chiasmolithus expansus Bramlette and Sullivan 1961

†Chiasmolithus grandis Bramlette and Riedel 1954

†Chiasmolithus modestus Perch-Nielsen 1971

†Chiasmolithus oamaruensis Deflandre 1954

†Chiasmolithus solitus Bramlette and Sullivan 1961

G. †Clausicoccus Prins 1979

G. Coronocyclus Hay et al. 1966

G. †Ericsonia Black 1964

G. †Gephyrorhabdus Hill 1976

G. †Holodiscolithus Roth 1970

†Holodiscolithus solidus Deflandre and Fert 1954

G. †Hughesius Varol 1989

G. †Praeprinsius Varol and Jakubowski 1989

G. †Solidopons Theodoridis 1984

Invalid names: Hymenomonadaceae Senn 1900 [empty], Pleurochrysidaceae Fresnel and Billard 1991 [empty]

G. †Daktylethra Gartner and Bukry 1969

Or. †Discoasterales Hay 1977

Fm. †Fasciculithaceae Hay and Mohler 1967

G. †Fasciculithus Bramlette and Sullivan 1961

G. †Lithoptychius Aubry 2011

Fm. †Heliolithaceae Hay and Mohler 1967

G. †Bomolithus Roth 1973

G. †Heliolithus Bramlette and Sullivan 1961

Invalid names: Discoasteraceae Tan Sin Hok 1927 [empty]

Fm. †Ellipsagelosphaeraceae Noël 1965

Subfm. †Actinosphaeroideae Noël 1965

G. †Calolithus Noël 1965

G. †Ellipsolithus Sullivan 1964

G. Gladiolithus Jordan and Chamberlain 1992

G. †Ilselithina Stradner and Adamiker 1966

Or. Isochrysidales Pascher 1910

Fm. Gephyrocapsaceae Heimdal 1997

G. Crenalithus Roth 1973

Fm. Noelaerhabdaceae Jerkovic 1970

G. Apertapetra Hay et al. 1966

G. Cepekiella Roth 1970

G. †Cribrocentrum Perch-Nielsen 1971

G. Dictyococcites Black 1967

†Dictyococcites scrippsae Bukry and Percival 1973

G. Emiliania Hay and Mohler 1967

G. Gephyrocapsa Kamptner 1943

G. †Pseudoemiliania Gartner 1969

Fm. †Prinsiaceae Hay and Mohler 1967

G. †Futyania Varol 1989

G. †Girgisia Varol 1989

G. Hornibrookina Edwards 1973

†Hornibrookina teuriensis Edwards 1973

†Hornibrookina weimerae Self-Trail 2011

G. †Neobiscutum Varol 1989

G. †Prinsius Hay and Mohler 1967

†Prinsius dimorphosus Perch-Nielsen 1977

†Prinsius martinii Haq 1971

†Prinsius tenuiculus Perch-Nielsen 1984

Invalid names: Isochrysidaceae Pascher 1910 [empty]

G. †Lanternithus Stradner 1962

†Lanternithus minutus Stradner 1962

G. †Minylitha Bukry 1973

G. Neosphaera Lecal-Schlauder 1949

G. †Pedinocyclus Bukry and Bramlette 1971

Fm. †Rhomboasteraceae Bown 2005

G. †Rhomboaster Bramlette and Sullivan 1961

G. †Tribrachiatus Shamrai 1963

G. Tetralithoides Theodoridis 1984

G. †Transversopontis Hay et al. 1966

Or. Zygodiscales Young and Bown 1997

Fm. Pontosphaeraceae Lemmermann 1908

G. Calciopilleus Müller 1974

G. Scyphosphaera Lohmann 1902

Fm. Zygodiscaceae Hay and Mohler 1967

G. †Isthmolithus Deflandre and Fert 1954

†Isthmolithus recurvus Deflandre 1954

G. †Lophodolithus Deflandre and Fert 1954

†Lophodolithus mochlophorus Deflandre 1954

G. †Nannotetrina Achuthan and Stradner 1969

G. †Neococcolithes Sujkowski 1931

†Neococcolithes dubius Deflandre and Fert 1954

†Neococcolithes protenus Black 1967

G. †Zygodiscus Bramlette and Sullivan 1961

Invalid names: Helicosphaeraceae Black 1971 [empty]

Invalid names: Phaeocystales Medlin 2000 [empty], Prymnesiales Papenfuss 1955 [empty], Prymnesiophyceae Hibberd 1976 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

low Mg calcitep

Environment:

oceanic, oligotrophic, marine, brackishp

Locomotion:

passively mobilep

Life habit:

planktonicp

Depth habitat:

surfacep

Diet:

"photoautotroph"p

Created:

2006-10-31 06:24:05

Modified:

2006-10-31 08:24:05

Source: p = phylum

Reference: Kiessling 2003

Age range

Maximum range based only on fossils: base of the Hettangian to the top of the Holocene or 201.40000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 199.5 Ma

Collections (10512 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Middle Hettangian - Late/Upper Hettangian	201.4 - 199.5	Canada (Alberta)	Tubirhabdus patulus, Crepidolithus crassus, Mitrolithus jansae, Mitrolithus elegans, Parhabdolithus liasicus, Crucirhabdus primulus (197630)
Hettangian - Sinemurian	201.4 - 192.9	Austria	Schizosphaerella punctulata (38056 38057 38058 38059)
Late/Upper Sinemurian - Late/Upper Pliensbachian	199.5 - 184.2	Morocco	Calyculus sp., Similiscutum cruciulus, Crepidolithus crassus, Schizosphaerella spp., Schizosphaerella sp., Calcivascularis jansae, Parhabdolithus robustus, Parhabdolithus liasicus, Crucirhabdus primulus (205771) Calyculus sp., Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella sp., Mitrolithus lenticularis, Mitrolithus elegans, Crucirhabdus primulus (205769) Lotharingius frodoi, Lotharingius haufii, Lotharingius sigillatus, Bussonius prinsii, Calyculus sp., Similiscutum finchii, Schizosphaerella sp., Schizosphaerella spp., Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus liasicus, Crucirhabdus primulus (205779) Lotharingius frodoi, Lotharingius haufii, Similiscutum finchii, Similiscutum cruciulus, Calcivascularis jansae, Mitrolithus elegans, Parhabdolithus liasicus (205774) Lotharingius frodoi, Similiscutum cruciulus, Schizosphaerella sp., Schizosphaerella spp., Calcivascularis jansae, Mitrolithus elegans, Parhabdolithus robustus, Parhabdolithus liasicus (205772) Lotharingius haufii, Schizosphaerella sp., Mitrolithus elegans (205776) Lotharingius haufii, Similiscutum novum, Similiscutum finchii, Crepidolithus crassus, Schizosphaerella sp., Schizosphaerella spp., Calcivascularis jansae (205778) Schizosphaerella sp. (205759 205760 205762 205763 205767 205768 205770 205775) Schizosphaerella sp., Calcivascularis jansae (205765) Schizosphaerella sp., Mitrolithus lenticularis (205764) Schizosphaerella sp., Schizosphaerella spp. (205761 205773) Schizosphaerella sp., Schizosphaerella spp., Calcivascularis jansae (205777)
Pliensbachian	192.9 - 184.2	Spain (Jaén)	Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus (201758) Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus (201759) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius barozii, Bussonius prinsii, Calyculus recondita, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus (201756) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius barozii, Bussonius prinsii, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201752) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Bussonius prinsii, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis (201754) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201753) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Mitrolithus elegans, Parhabdolithus distinctus, Crucirhabdus primulus (201755)
Spinatum	192.9 - 184.2	Morocco	Lotharingius sigillatus (203735 203736)
Early/Lower Toarcian	184.2 - 182.9	Spain (Granada)	Watznaueria colacicchii, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus superbus, Discorhabdus striatus, Similiscutum sp. 1, Biscutum intermedium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199494)
Toarcian	184.2 - 174.7	Spain (Jaén)	Lotharingius sigillatus, Lotharingius hauffii, Lotharingius barozii, Bussonius prinsii, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201678) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius barozii, Bussonius prinsii, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201750) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius barozii, Bussonius prinsii, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201675) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius barozii, Bussonius prinsii, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus (201751) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Bussonius prinsii, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201672) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Bussonius prinsii, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201676) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Bussonius prinsii, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201665) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201671) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Mitrolithus elegans, Parhabdolithus distinctus (201667) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201670) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus (201677) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus (201685) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus recondita, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201673) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201674) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus (201666) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Tubirhabdus patulus, Crepidolithus cavus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201668) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crucifer, Crepidolithus granulatus, Crepidolithus crassus, Schizosphaerella punctulata, Diductius constans, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus (201669)
Toarcian	184.2 - 174.7	Saudi Arabia	Ellipsagelosphaera britannica, Tetralithus gothicus (45089) Ellipsagelosphaera britannica, Tetralithus goticus (45191)
Toarcian	184.2 - 174.7	Spain (Murcia)	Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (201778) Lotharingius hauffii, Carinolithus superbus, Discorhabdus striatus, Schizosphaerella punctulata (201774) Lotharingius hauffii, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201781) Lotharingius sigillatus, Carinolithus superbus, Discorhabdus striatus, Biscutum intermedium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201787) Lotharingius sigillatus, Carinolithus superbus, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (201792) Lotharingius sigillatus, Lotharingius hauffii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Crepidolithus crassus, Schizosphaerella punctulata (201772) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Crepidolithus crassus, Schizosphaerella punctulata (201776) Watznaueria colacicchii, Lotharingius sigillatus, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201788) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201764) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201780) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Biscutum dubium, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (201771) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata (201779) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201794) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata (201775) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata (201765 201766) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201793) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (201768) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201777) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius velatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201782) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Biscutum dubium, Triscutum sullivanii, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201773) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201783) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (201790) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Triscutum sullivanii, Crepidolithus crassus, Schizosphaerella punctulata (201770) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius barozii, Lotharingius velatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201791) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Triscutum sullivanii, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201789) Watznaueria colacicchii, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201769)
Middle Toarcian	182.9 - 180.4	Spain (Murcia)	Bussonius prinsii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Biscutum dubium, Crepidolithus crassus (201809) Lotharingius hauffii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata, Mitrolithus lenticularis (201808) Lotharingius hauffii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201811) Lotharingius hauffii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201810) Lotharingius hauffii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (201805) Lotharingius hauffii, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata (201807) Lotharingius hauffii, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus superbus, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (201806) Lotharingius sigillatus, Lotharingius hauffii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata, Mitrolithus lenticularis (201803) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata (201799) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Crepidolithus crassus, Schizosphaerella punctulata (201813) Lotharingius sigillatus, Lotharingius hauffii, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201812) Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata, Mitrolithus lenticularis (201802) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201795) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Bussonius prinsii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Discorhabdus cf. D. criotus, Discorhabdus striatus, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata (201804) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata, Mitrolithus lenticularis (201798) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata, Mitrolithus lenticularis (201800) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201801) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201797) Watznaueria sp. A, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Bussonius prinsii, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata, Mitrolithus lenticularis (201796)
Levesquei	182.9 - 174.7	Germany	Bussonius leufuensis (59428) Lotharingius crucicentralis, Carinolithus superbus, Triscutum sullivanii, Retecapsa incompta, Crepidolithus crassus, Crucirhabdus primulus (59434) Watznaueria contracta, Lotharingius crucicentralis, Lotharingius velatus, Bussonius leufuensis, Carinolithus crassus, Carinolithus superbus, Discorhabdus criotus, Triscutum tiziense, Triscutum sullivanii, Retecapsa incompta, Crepidolithus crassus, Schizosphaerella sp., Crucirhabdus primulus (59430) Watznaueria contracta, Lotharingius velatus, Discorhabdus criotus, Retecapsa incompta, Schizosphaerella sp. (59431)
Late/Upper Toarcian	180.4 - 174.7	Spain (Granada)	Carinolithus cantaluppii, Carinolithus poulnabronei, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (202110) Carinolithus cantaluppii, Carinolithus poulnabronei, Discorhabdus striatus, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (202114) Carinolithus cantaluppii, Discorhabdus striatus, Similiscutum novum, Crepidolithus crassus, Schizosphaerella punctulata (202108) Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (202103) Carinolithus poulnabronei, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Crepidolithus crassus, Schizosphaerella punctulata (202117) Crepidolithus crassus, Schizosphaerella punctulata (202113) Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (202106) Lotharingius crucicentralis, Lotharingius velatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Crepidolithus crassus, Schizosphaerella punctulata (199235) Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Discorhabdus sp. cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum intermedium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (202097) Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Crepidolithus crassus, Schizosphaerella punctulata (202100) Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199496) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum intermedium, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199177) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (202107) Lotharingius sigillatus, Lotharingius velatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (202098) Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius velatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus sp. cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (202101) Lotharingius velatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Similiscutum novum, Crepidolithus crassus, Schizosphaerella punctulata (202116) Lotharingius velatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (202111) Watznaueria colacicchii, Lotharingius sigillatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (202115) Watznaueria colacicchii, Lotharingius sigillatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Crepidolithus crassus, Schizosphaerella punctulata (202105) Watznaueria colacicchii, Lotharingius sigillatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199198) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199487) Watznaueria colacicchii, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199248) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum intermedium, Triscutum sullivanii, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199495) Watznaueria colacicchii, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199253) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Triscutum sullivanii, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199165) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum depravatum, Crepidolithus crassus, Schizosphaerella punctulata (201880) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (201948) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Bussonius prinsii, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199169) Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199164) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199493) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199168) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199229) Watznaueria colacicchii, Watznaueria sp. A, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199195) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum intermedium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199187) Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Triscutum sullivanii, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199171) Watznaueria colacicchii, Watznaueria sp. B, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum sp. 1, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199492) Watznaueria colacicchii, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Similiscutum novum, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199173) Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum depravatum, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199167) Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Biscutum depravatum, Biscutum intermedium, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199166) Watznaueria sp. A, Discorhabdus striatus, Similiscutum novum, Crepidolithus crassus, Schizosphaerella punctulata (202104) Watznaueria sp. A, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Crepidolithus crassus, Schizosphaerella punctulata (202109) Watznaueria sp. A, Lotharingius sigillatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (202099) Watznaueria sp. A, Lotharingius sigillatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Discorhabdus striatus, Crepidolithus crassus, Schizosphaerella punctulata (202112) Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum intermedium, Crepidolithus crassus, Schizosphaerella punctulata (199181) Watznaueria sp. A, Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus sp. cf. D. criotus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum sp. 1, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199490) Watznaueria sp. A, Watznaueria sp. B, Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum intermedium, Biscutum dubium, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199247) Watznaueria sp. B, Carinolithus poulnabronei, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Schizosphaerella punctulata (202102) Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Biscutum dubium, Crepidolithus crassus, Schizosphaerella punctulata (199231) Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199491) Watznaueria sp. B, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Biscutum intermedium, Tubirhabdus patulus, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata (199488) Watznaueria sp. B, Lotharingius sigillatus, Lotharingius hauffii, Lotharingius crucicentralis, Lotharingius velatus, Bussonius prinsii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus ignotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum dubium, Tubirhabdus patulus, Crepidolithus crassus, Schizosphaerella punctulata (199172)
Late/Upper Toarcian	180.4 - 174.7	Germany	Carinolithus superbus, Discorhabdus striatus, Retecapsa incompta, Crepidolithus crassus, Schizosphaerella sp. (59454) Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius contractus, Lotharingius barozii, Bussonius sp. 1, Bussonius prinsii, Bussonius leufuensis, Carinolithus superbus, Calyculus sp. 1, Calyculus sp. 2, Proculithus fistulatus, Catillus hommerilii, Calyculus cribrum, Discorhabdus novus, Biscutum striatum, Palaeopontosphaera dubia, Biscutum intermedium, Axopodorhabdus atavus, Axopodorhabdus cylindratus, Tubirhabdus patulus, Tubirhabdus patulus, Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Crepidolithus crassus, Vagalapilla quadriarculla, Schizosphaerella punctulata, Parhabdolithus cavus, Stradnerlithus humilis (59451) Lotharingius haufforum, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius barozii, Bussonius sp. 1, Bussonius prinsii, Bussonius leufuensis, Carinolithus sceptrum, Carinolithus superbus, Proculithus charlottei, Catillus hommerilii, Calyculus cribrum, Discorhabdus novus, Biscutum criotum, Biscutum sp. 2, Biscutum sp. 4, Biscutum striatum, Palaeopontosphaera dubia, Biscutum intermedium, Axopodorhabdus depravatus, Axopodorhabdus atavus, Axopodorhabdus cylindratus, Ethmorhabdus crucifer, Triscutum beaminsterensene, Triscutum sullivanii, Tubirhabdus patulus, Tubirhabdus patulus, Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata, Parhabdolithus cavus, Crucirhabdus magnus, Stradnerlithus humilis (59453) Lotharingius sp. 2, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius contractus, Lotharingius barozii, Bussonius sp. 1, Bussonius prinsii, Bussonius leufuensis, Carinolithus sp. 1, Carinolithus sceptrum, Carinolithus superbus, Calyculus sp. 2, Proculithus fistulatus, Proculithus charlottei, Discorhabdus novus, Biscutum criotum, Biscutum striatum, Palaeopontosphaera sp. 1, Palaeopontosphaera dubia, Biscutum intermedium, Axopodorhabdus depravatus, Axopodorhabdus atavus, Axopodorhabdus cylindratus, Triscutum pricatillus, Tubirhabdus patulus, Zeugrhabdotus choffatii, Zeugrhabdotus sp. 1, Zeugrhabdotus sp. 2, Zeugrhabdotus erectus, Crepidolithus crassus, Vagalapilla quadriarculla, Schizosphaerella punctulata, Parhabdolithus cavus, Crucirhabdus magnus (59449)
Early/Lower Aalenian	174.7 - 170.9	Germany	Bussonius prinsii, Calyculus sp. (59458) Bussonius prinsii, Carinolithus superbus, Calyculus sp., Discorhabdus striatus, Retecapsa incompta, Crepidolithus crassus, Schizosphaerella sp. (59457)
Bajocian	170.9 - 168.2	Saudi Arabia	Cyclagelosphaera margereli (45268) Ellipsagelosphaera britannica, Schizosphaerella punctulata, Tetralithus gothicus (45265)
Bajocian	170.9 - 168.2	Spain	Cyclagelosphaera margerelii, Watznaueria communis, Watznaueria contracta, Watznaueria britannica (214379)
Bathonian	168.2 - 165.3	Saudi Arabia	Cyclagelosphaera margereli, Ellipsagelosphaera britannica, Diazomatolithus lehmani, Tetralithus gothicus (45519) Cyclagelosphaera margereli, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Hexalithus magharensis, Schizosphaerella punctulata, Tetralithus gothicus (45464)
Callovian	165.3 - 161.5	Saudi Arabia	Cyclagelosphaera margereli, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Tetralithus gothicus (45549) Cyclagelosphaera margereli, Ellipsagelosphaera communis (45555) Cyclagelosphaera margereli, Tetralithus gothicus (45558) Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis (45540) Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Discorhabdus sp., Ethmorhabdus gallicus, Zeugrhabdotus erectus, Stephanolithion bigoti, Tetralithus gothicus, Zygodiscus sp. (45541) Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Diazomatolithus lehmani, Tetralithus gothicus (45547) Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Ethmorhabdus anglicus, Polypodorhabdus escaigii, Zeugrhabdotus erectus, Stephanolithion bigoti (47890)
Middle Oxfordian	161.5 - 154.8	Switzerland	Cyclagelosphaera margereti, Ellipsagelosphaera britannica, Discorhabdus patulus, Axopodorhabdus cylindratus, Ethmorhabdus gallicus, Hexapodorhabdus cuvillieri, Crepidolithus perforatus, Vekshinella quadriarculla, Stephanolithion bigoti (58568)
Oxfordian	161.5 - 154.8	Saudi Arabia	Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera communis (46052) Watznaueria manivitae, Biscutum sp., Ethmorhabdus sp., Vekshinella stradneri (45896)