PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Groebertherium allenensis

Mammalia - Dryolestidae

Taxonomy

Barberenia allenensis was named by Rougier et al. (2009). Its type specimen is MML-Pv 13, a tooth (right upper molariform), and it is a 3D body fossil. Its type locality is Cerro Tortuga, Bajo Trapalcó, which is in a Campanian/Maastrichtian fluvial sandstone/siltstone in the Allen Formation of Argentina.

It was recombined as Groebertherium allenensis by Connelly et al. (2024).

Year	Name and author
2009	Barberenia allenensis Rougier et al. p. 230 figs. 7-8
2024	Groebertherium allenensis Connelly et al.

Reference	Diagnosis
G. W. Rougier et al. 2009	Medium sized dryolestoid with a skewed compressed basin similar to Quirogatherium and Barberenia araujoae (Bonaparte, 1990). Differs from Quirogatherium in the more gracile cusp pattern, smaller size and more skewed cusp arrangement. Barberenia allenensis is similar to B. araujoae in the sharp, tall cusps, but the Allen Formation species is distinctly larger, has a less mesiodistally elongated triangular outline with a smaller parastylar area and a less developed anterior cingulum. The parastylar hook is relatively small, signiﬁcantly smaller than the similar Barberenia araujoae (Bonaparte, 1990).
B. E. Connelly et al. 2024	Small-sized dryolestoid, slightly smaller than G. stipanicici yet with more robust and bulging cusps. It has a skewed compressed basin, similar to Quirogatherium major and G. stipanicici. Differs from Quirogatherium 448 in the more gracile cusp pattern, smaller size, and more skewed cusp arrangement. The holotype of G. allenensis is similar to G. stipanicici (MACN-RN 1034) in having tall, sharp cusps, but the Allen Formation premolar has a less mesio-distally elongated triangular outline with smaller parastylar area and parastylar hook, and less developed anterior cingulum. All specimens here referred to G. allenensis differ from G. stipanicici in having a less pronounced ectoflexus. They also exhibit a reduced metastylar area, which does not protrude as in G. stipanicici but is nearly aligned with the postparacrista.