PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Cypriniformes (carp-like fish)

Actinopteri - Cypriniformes

Taxonomy

Cypriniformes was named by Bleeker (1859). It is extant.

It was assigned to Osteichthyes by Mors et al. (2000); to Ostariophysi by Nelson (2006); to Actinopterygii by Böhme (2010); to Otophysi by Long (2011); to Teleostei by Böhme (2008) and Fierstine et al. (2012); to Cyprinae by Betancur et al. (2013); and to Cypriniphysi by Nelson et al. (2016).

Subtaxa

Botiidae, Catostomidae, Cobitidae, Cobitoidea, Cyprinidae, Cyprinoidea, Gyrinocheilidae, Nemacheilidae, Siluri

Synonymy list

Year	Name and author
1859	Cypriniformes Bleeker
2000	Cypriniformes Mors et al. p. 151
2006	Cypriniformes Nelson
2008	Cypriniformes Böhme p. 163
2010	Cypriniformes Böhme p. 6
2011	Cypriniformes Long p. 243
2012	Cypriniformes Fierstine et al. p. 30
2013	Cypriniformes Betancur et al.
2016	Cypriniformes Nelson et al. p. 180

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
superclass	Actinopterygii	()
class	Actinopteri	()
subclass	Neopterygii	(Regan 1923)

Rank	Name	Author
—	Teleosteomorpha
—	Teleostei	(Müller 1846)
—	Teleocephala	de Pinna 1996
—	Clupeocephala	Patterson and Rosen 1977
—	Otomorpha
—	Ostariophysi	(Jordan and Evermann 1896)
—	Otophysa
superorder	Cyprinae
order	Cypriniformes	Bleeker 1859
order	Cypriniformes	Bleeker 1859

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Cypriniformes Bleeker 1859 [carp-like fish]

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Fm. Catostomidae

G. Carpiodes Rafinesque 1820

Carpiodes cyprinus LeSueur 1817

G. †Catostomus

Catostomus batrachops Cope 1883

Catostomus catostomus Forster 1773

Catostomus commersonnii Lacépède 1803

†Catostomus cristatus Cope 1883

Catostomus labiatus Ayres 1855

Catostomus shoshonensis Cope 1883

G. †Chasmistes Jordan 1878

†Chasmistes cujus Cope 1883

Subfm. †Ictiobinae Alvarado-Ortega et al. 2006

G. †Ictiobus Alvarado-Ortega et al. 2006

†Ictiobus aguilerai Alvarado-Ortega et al. 2006

G. †Xyrauchen Eigenmann and Kirsch 1889

Xyrauchen texanus Abbott 1860

Invalid names: Catostomini [empty], Cycleptinae Nelson et al. 2016 [empty], Erimyzontini Nelson et al. 2016 [empty], Moxostomatini [empty], Myxocyprininae [empty], Thoburniini [empty]

Fm. Cobitidae

G. Cobitis Linnaeus 1758

†Cobitis centrochir Agassiz 1832

†Cobitis cephalotes Agassiz 1832

†Cobitis primigenius Böhme 2008

Invalid names: Acanthopsis Agassiz 1832 [synonym]

Superfm. Cobitoidea Swainson 1838

Invalid names: Balitoridae [empty], Barbuccidae [empty], Ellopostomatidae [empty], Gastromyzontidae [empty], Jianghanichthyidae [empty], Serpenticobitidae [empty], Vaillantellidae [empty]

Fm. Cyprinidae Rafinesque 1815

G. Acrocheilus Agassiz 1855

†Acrocheilus latus Cope 1870

Invalid names: Diastichus macrodon Cope 1870 [synonym]

G. †Alisodon Hay 1920

†Alisodon mirus Hay 1920

G. †Amblypharyngodon Bleeker 1860

G. †Amyzon Cope 1872

†Amyzon brevipinne Cope 1893

†Amyzon commune Cope 1874

†Amyzon fusiforme Cope 1875

†Amyzon gosiutensis Grande et al. 1982

†Amyzon mentale Cope 1872

†Amyzon pandatum Cope 1875

G. Aspius Agassiz 1832

†Aspius brongniarti Agassiz 1835

†Aspius ecnomi Sauvage 1870

†Aspius gracilis Agassiz 1835

G. Barbus Cuvier and Cloquet 1816

Barbus altianalis Boulenger 1900 [ripon barbel]

Barbus barbus Linnaeus 1758

Barbus bynni Forsskål 1775 [binny]

Subfm. Cyprininae Bonaparte 1840

G. †Aphelichthys Cope 1893

†Aphelichthys lindahlii Cope 1893

G. †Diastichus Cope 1870

†Diastichus parvidens Cope 1870

†Diastichus strangulatus Cope 1870

G. Notropis Rafinesque 1818

†Notropis angustarcus Cope 1877

G. †Palaeocarassius Obrhelová 1970

G. Rutilus Rafinesque 1820

†Rutilus altarcus Cope 1877

†Rutilus condonianus Cope 1883

†Rutilus gibbarcus Cope 1877

Invalid names: Anchybopsis Cope 1870 [synonym]

G. Cyprinion Heckel 1843

Cyprinion macrostomum Heckel 1843

G. Cyprinus Linnaeus 1758

G. †Ecocarpia Chen et al. 2005

†Ecocarpia ningmingensis Chen et al. 2005

G. †Gila Baird and Girard 1853

†Gila coerulea Girard 1856

†Gila milleri Smith 1975

G. †Gobio Cuvier 1817

†Gobio analis Agassiz 1835

G. †Hexapsephus Gunther 1876

†Hexapsephus guentheri Woodward 1901

G. †Idadon Smith 1975

†Idadon condonianus Smith 1975

†Idadon hibbardi Smith 1975

G. †Iquius Jordan 1919

†Iquius nipponicus Jordan 1919

Subfm. Labeoninae Chen 1984

Tr. Garraini Reid 1982

G. Garra Hamilton 1822

G. †Lavinia Girard 1845

G. †Leuciscus Cuvier 1817

Subg. †Leuciscus (Scardinius) Sauvage 1873

†Leuciscus (Scardinius) dumerilii Sauvage 1870

†Leuciscus (Scardinius) sicanus Sauvage 1873

Subg. Leuciscus (Squalius) Bonaparte 1837

†Leuciscus arciferus Cope 1870

†Leuciscus bairdii Cope 1870

†Leuciscus brevis Agassiz 1839

†Leuciscus cephalon Zenker 1833

†Leuciscus gracilis Agassiz 1839

†Leuciscus hartmanni Agassiz 1839

†Leuciscus heterurus Agassiz 1832

†Leuciscus laminatus Cope 1870

†Leuciscus latiusculus Agassiz 1839

†Leuciscus leptus Agassiz 1832

†Leuciscus oeningensis Agassiz 1832

†Leuciscus poeticus Cope 1870

†Leuciscus pusillus Agassiz 1835

†Leuciscus reddingi Cope 1883

†Leuciscus turneri Lucas 1900

Invalid names: Oligobelus Cope 1870 [synonym], Squalius Bonaparte 1837 [synonym]

G. †Mylocyprinus Leidy 1870

†Mylocyprinus inflexus Cope 1883

†Mylocyprinus robustus Leidy 1870

G. Nemachilus Günther 1868

G. Orthodon Girard 1856

†Orthodon condamyi Coquand 1859

†Orthodon hadrognathus Smith 1975

G. †Palaeoleuciscus Obrhelová 1969

†Leuciscus (Palaeoleuciscus) soccoloviensis Obrhelova 1969

†Palaeoleuciscus chartaceus Laube 1905

G. †Palaeotinca Obrhelová 1969

†Palaeotinca egeriana Obrhelová 1969

Subfm. Phoxininae

G. †Palaeorutilus Gaudant 1988

†Palaeorutilus medius Reuss 1844

Invalid names: Leuciscus (Palaeoleuciscus) luzicensis Obrhelovd 1969 [synonym]

†Palaeorutilus papyraceus Bronn 1828

G. †Planktophaga Böhme et al. 2013

†Planktophaga minuta Böhme et al. 2013

G. †Protothymallus Laube 1901

†Protothymallus elongata Kramberger 1884

Invalid names: Varhostichthys Obrhelová 1969 [synonym]

G. †Ptychocheilus Agassiz 1855

†Ptychocheilus arciferus Cope 1870

G. Puntius Hamilton 1822

G. †Rhodeus Agassiz 1832

†Rhodeus elongatus Agassiz 1835

†Rhodeus latior Agassiz 1835

G. †Siphateles Cope 1883

†Siphateles bicolor Girard 1856

†Siphateles mohavensis Snyder 1918

G. †Thynichthys Bleeker 1860

G. †Tinca Cuvier 1817

†Tinca furcata Agassiz 1832

†Tinca leptosoma Agassiz 1832

†Tinca micropygoptera Agassiz 1839

Subfm. Tincinae

G. †Tarsichthys Troschel 1854

†Tarsichthys macrurus Agassiz 1839

Invalid names: Tinca tarsiger Troschel 1854 [synonym]

Invalid names: Acheilognathinae [empty], Barbinae [empty], Cultrinae [empty], Danioninae [empty], Esox destructus Laube 1901 [nomen vanum], Gobioninae Bleeker 1863 [empty], Leuciscinae Howes 1991 [empty], Opsariichthyinae [empty], Squaliobarbinae [empty], Xenocyprinae [empty]

Superfm. Cyprinoidea

Invalid names: Psilorhynchidae Hora 1926 [empty]

Invalid names: Botiidae [empty], Gyrinocheilidae [empty], Nemacheilidae [empty], Siluri Stirton 1953 [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

freshwatero

Locomotion:

actively mobileo

Life habit:

nektonico

Diet:

omnivoreo

Created:

2009-07-20 20:40:14

Modified:

2017-04-17 10:32:02

Source: o = order, subp = subphylum

References: Hendy et al. 2009, Carroll 1988

Age range

Maximum range based only on fossils: base of the Late/Upper Tithonian to the top of the Holocene or 150.80000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 145.0 Ma

Collections (231 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Tithonian	152.1 - 145.0	Greenland	Cyprinidae indet. (1212 1214)
Late/Upper Tithonian - Early/Lower Berriasian	150.8 - 140.2	Morocco (Oriental)	Cypriniformes indet. (45726)
Ypresian	56.0 - 47.8	Canada (British Columbia)	Amyzon brevipinne (144348 204404)
Eocene	56.0 - 33.9	Canada (British Columbia)	Amyzon sp. (32896)
Eocene - Early/Lower Oligocene	56.0 - 28.4	USA (Oregon)	Catostomidae indet., Amyzon sp. (227073)
Lutetian	47.8 - 41.3	Pakistan (North-West Frontier)	Cyprinidae indet. (28978)
Lutetian	47.8 - 41.3	USA (Washington)	Amyzon sp. (189961)
Lutetian	47.8 - 41.3	Canada (British Columbia)	Amyzon sp. (189959)
Lutetian	47.8 - 41.3	USA (Wyoming)	Amyzon gosiutensis (192667 192668)
Lutetian - Bartonian	47.8 - 38.0	China (Guangdong)	Cyprinus maomingensis (167230)
Early/Lower Uintan	46.2 - 39.7	USA (Colorado)	Catostomidae indet. (16623)
Bartonian - Late/Upper Eocene	41.3 - 33.9	Vietnam (Lang Son)	Planktophaga minuta, Barbinae indet., Xenocyprinae indet., Cultrinae indet., Acheilognathidae indet. (169422)
Bartonian - Priabonian	41.3 - 33.9	Vietnam (Lang Son)	Barbinae indet. (169420)
Duchesnean	39.7 - 37.0	USA (Utah)	Catostomidae indet. (16877)
Late/Upper Eocene	37.2 - 33.9	China (Guangdong)	Cyprinus maomingensis (144065)
Early/Lower Oligocene	33.9 - 28.4	China (Guangxi)	Cobitis nanningensis (228111 228112)
MP 21	33.9 - 28.4	Czech Republic	Leuciscus colei, Leuciscus stephani, Leuciscus brevis (184466)
MP 23	33.9 - 28.4	Switzerland (Canton Jura)	Leuciscus sp. (183544)
Rupelian	33.9 - 28.1	Romania	Barbus sp., Scardinius sp. (50344)
Oligocene	33.9 - 23.03	Czech Republic (Tschechien)	Cyprinidae indet. (185664)