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Clupeomorpha
Taxonomy
Clupeomorpha was named by Greenwood et al. (1966). It is extant.
It was reranked as the superorder Clupeomorpha by Murray et al. (2005); it was reranked as the unranked clade Clupeomorpha by De Figueiredo (2009), Bannikov (2014).
It was assigned to Teleostei by Nelson (2006); to Ostarioclupeomorpha by Long (2011); to Teleostei by Forey et al. (2003), De Figueiredo (2009), Bannikov (2014), Marramá and Carnevale (2015); to Otocephala by Nelson et al. (2016); and to Clupeocephala by Murray et al. (2005), Than-Marchese et al. (2024).
It was reranked as the superorder Clupeomorpha by Murray et al. (2005); it was reranked as the unranked clade Clupeomorpha by De Figueiredo (2009), Bannikov (2014).
It was assigned to Teleostei by Nelson (2006); to Ostarioclupeomorpha by Long (2011); to Teleostei by Forey et al. (2003), De Figueiredo (2009), Bannikov (2014), Marramá and Carnevale (2015); to Otocephala by Nelson et al. (2016); and to Clupeocephala by Murray et al. (2005), Than-Marchese et al. (2024).
Synonymy list
| Year | Name and author |
|---|---|
| 1966 | Clupeomorpha Greenwood et al. p. 350 |
| 2003 | Clupeomorpha Forey et al. p. 229 |
| 2005 | Clupeomorpha Murray et al. p. 301 |
| 2006 | Clupeomorpha Nelson |
| 2009 | Clupeomorpha De Figueiredo |
| 2011 | Clupeomorpha Long p. 243 |
| 2014 | Clupeomorpha Bannikov p. 25 |
| 2015 | Clupeomorpha Marramá and Carnevale |
| 2016 | Clupeomorpha Nelson et al. p. 162 |
| 2024 | Clupeomorpha Than-Marchese et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Superor. Clupeomorpha Greenwood et al. 1966
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Or. †Ellimmichthyiformes Grande 1982
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Fm. †Armigatidae Murray and Wilson 2013
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G. †Armigatus Grande 1982
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†Armigatus alticorpus Forey et al. 2003
†Armigatus brevissimus de Blainville 1818
†Armigatus carrenoae Alvarado-Ortega et al. 2020
†Armigatus dalmaticus Murray et al. 2016
†Armigatus elatus Costa 1850
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Invalid names: Histiurus serioloides Costa 1864 [synonym], Histiurus ventricosus Costa 1865 [synonym]
†Armigatus namourensis Forey et al. 2003
†Armigatus oligodentatus Vernygora and Murray 2015
†Armigatus plinii Marramà and Carnevale 2023
Invalid names: Histiurus Costa 1850 [nomen oblitum]
Subor. †Armigatoidei Murray and Wilson 2013
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G. †Diplomystus Cope 1877
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†Diplomystus altisomus Yabumoto 1994
†Diplomystus altus Leidy 1873
†Diplomystus birdi Woodward 1895
†Diplomystus coverhamensis Chapman 1918
†Diplomystus dartevellei Casier 1965
†Diplomystus dentatus Cope 1877
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Invalid names: Diplomystus analis Cope 1877 [synonym], Diplomystus pectorosus Cope 1877 [synonym], Diplomystus theta Cope 1873 [synonym]
†Diplomystus dubertreti Signeux 1951
†Diplomystus kokuraensis Uyeno 1979
†Diplomystus primotinus Uyeno 1979
†Diplomystus vectensis Newton 1889
Invalid names: Copeichthys Dollo 1904 [objective synonym]
Fm. †Paraclupeidae Chang and Chou 1977
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G. †Ellimichthys Jordan and Gilbert 1919
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†Ellimmichthys goodi Eastman 1912
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Invalid names: Diplomystus elberti Weiler 1922 [synonym], Ellimma guineensis Gayet 1989 [synonym]
†Ellimmichthys longicostatum Cope 1880
†Ellimmichthys longicostatus Cope 1885
Subfm. †Paraclupeinae Chang and Chou 1977
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G. †Paraclupea Sun 1956
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†Paraclupea pietrarojae Marramà and Carnevale 2023
†Paraclupea seilacheri Alvarado-Ortega and Melgarejo-Damián 2017
G. †Scutatoclupea Bannikov 2015
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†Scutatoclupea applegatei Alvarado-Ortega and Ovalles-Damián 2008
†Scutatoclupea bacchiai Bannikov 2015
G. †Scutatuspinosus Silva Santos and Correa 1985
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†Scutatuspinosus itapagipensis Silva Santos and Correa 1985
G. †Thorectichthys Murray and Wilson 2013
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†Thorectichthys fideli Than-Marchese et al. 2024
†Thorectichthys marocensis Murray and Wilson 2013
†Thorectichthys rhadinus Murray and Wilson 2013
G. †Triplomystus Forey et al. 2003
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†Triplomystus noorae Forey et al. 2003
†Triplomystus oligoscutatus Forey et al. 2003
Invalid names: Ellimichthyidae Grande 1982 [synonym], Ellimmichthyinae [empty], Ellimminae [empty], Scutatuspinosinae Santos and Corréa 1985 [invalid subgroup], Thorectichthyinae [empty]
Fm. †Sorbinichthyidae Bannikov and Bacchia 2000
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Invalid names: Ellimichthyoidei [empty]
G. †Scombroclupea Kner 1863
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†Scombroclupea diminuta Forey et al. 2003
†Scombroclupea gaudryi Pictet and Humbert 1866
†Scombroclupea javieri Than-Marchese et al. 2020
†Scombroclupea macrophthalma Heckel 1849
†Scombroclupea occidentalis Giersch et al. 2011
†Scombroclupea scutata Woodward 1908
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, subp = subphylum | |||||
| References: Kiessling 2004, Carroll 1988, Hendy et al. 2009 | |||||
Age range
Maximum range based only on fossils: base of the Valanginian to the top of the Wasatchian or 137.05000 to 50.50000 Ma
Minimum age of oldest fossil (stem group age): 125.77 Ma
Minimum age of oldest fossil (stem group age): 125.77 Ma
Collections (83 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Neocomian | Japan (Fukuoka) | Diplomystus primotinus, Diplomystus kokuraensis (144936 145245) Diplomystus sp., Diplomystus altisomus (145246) | |
| Valanginian - Barremian | Spain (Biscay) | Ezkutuberezi carmeni (144740) | |
| Barremian - Maastrichtian | China (Nei Mongol) | Lycoptera sp. (91404) | |
| Aptian | Mexico (Chiapas) | Ellimmichthyiformes indet., Triplomystus applegatei, Paraclupea sp. (68835) | |
| Aptian - Albian | Equatorial Guinea | Diplomystus sp., Diplomystus goodi (129869) | |
| Early/Lower Albian | Italy (Campania) | Diplomystus sp. (68888 112631) | |
| Early/Lower Albian | Italy (Benevento) | Histiurus serioloides, Histiurus elatus, Histiurus ventricosus, Armigatus plinii, Paraclupea pietrarojae (127350) Paraclupea pietrarojae (237947) | |
| Albian | Mexico (Chiapas) | Thorectichthys sp. (241349) | |
| Albian - Cenomanian | Mexico (Hidalgo) | Clupeomorpha indet. (135815) | |
| Middle Albian | Canada (Northwest Territories) | Foreyclupea loonensis (94490) | |
| Middle Albian - Late/Upper Albian | Mexico (Puebla) | Ellimmichthys sp., Ranulfoichthys dorsonudum, Armigatus carrenoae (80283) Paraclupea seilacheri (133991 221635 221636 221637 221638) | |
| Cenomanian | Slovenia | Clupea gaudryi (144686) | |
| Cenomanian | New Zealand | Diplomystus coverhamensis (88982) | |
| Early/Lower Cenomanian | Palestine | Rhombichthys intoccabilis (11804) | |
| Early/Lower Cenomanian | Mexico (Chiapas) | Paraclupeidae indet., Scombroclupea javieri, Thorectichthys fideli (192074) | |
| Late/Upper Cenomanian | Lebanon | Armigatus alticorpus (221149) Clupea brevissima, Clupea sardinoides (221186) Clupea brevissima, Scombroclupea macrophthalma, Clupea sardinoides, Clupea gaudryi (221227) Clupea gaudryi (69118) Clupea macrophthalma (221190) Clupea sardinoides (221228) Ornategulum sardinioides (221948) | |
| Late/Upper Cenomanian | Morocco (Er Rachida) | Ellimmichthys sp., Diplomystus sp. (120457) | |
| Late/Upper Cenomanian | Croatia | Clupea brevissima, Scombroclupea macrophthalma, Clupea gaudryi (123565) | |
| Late/Upper Cenomanian | Slovenia | Clupea gaudryi (221237) | |
| Late/Upper Cenomanian | Morocco (Meknes-Tafilalet) | Armigatus oligodentatus (136753) | |
| Late/Upper Cenomanian | Canada (Alberta) | Tycheroichthys dunveganensis (125734) | |
| Middle Cenomanian | Lebanon | Scutatoclupea bacchiai (221634) Sorbinichthys elusivo (221170 221171 221633) Triplomystus noorae, Triplomystus oligoscutatus, Armigatus namourensis, Armigatus alticorpus, Scombroclupea macrophthalma, Scombroclupea diminuta, Ornategulum sardinioides (124426) | |
| Middle Cenomanian - Late/Upper Cenomanian | USA (Utah) | Ellimmichthyiformes indet. LvD, Ellimmichthyiformes indet. U-7, Diplomystus sp. (14385) | |
| Middle Cenomanian - Late/Upper Cenomanian | Mexico (Coahuila) | Scombroclupea occidentalis (221656) | |
| Cenomanian - Turonian | United Kingdom (England) | Diplomystus sp. (238510) | |
| Turonian | Croatia | Armigatus dalmaticus (221146) | |
| Late/Upper Turonian - Early/Lower Coniacian | Argentina (Neuquén) | Clupeomorpha indet., Semionotidae indet. (49415) | |
| Late/Upper Santonian | USA (South Dakota) | Clupeomorpha indet. (194130) | |
| Late/Upper Santonian | Canada (Alberta) | Horseshoeichthys sp. (14425 14426) | |
| Santonian - Campanian | Bolivia | Gasteroclupea branisai (219931) | |
| Late/Upper Campanian | USA (Alaska) | Horseshoeichthys armaserratus (240913) | |
| Middle Campanian | Canada (Alberta) | Horseshoeichthys sp. (181024) | |
| Middle Campanian | USA (Kansas) | Clupeomorpha indet. (25692) | |
| Middle Campanian - Late/Upper Campanian | Canada (Saskatchewan) | Diplomystus sp., Horseshoeichthys sp. (229975) | |
| Early/Lower Maastrichtian | Canada (Alberta) | Sorbinichthyidae indet. (135741) Sorbinichthyidae indet., Horseshoeichthys armaserratus (55363) | |
| Late/Upper Maastrichtian | Bolivia (Cochabamba) | Gasteroclupea branisai (12855) | |
| Late/Upper Maastrichtian | Argentina (Salta) | Gasteroclupea branisai (114766) | |
| Maastrichtian - Danian | Bolivia | Clupeidae indet. (229878) Gasteroclupea branisai (94770 219905 219909 219925 219926 219929 219930 219940) | |
| Maastrichtian - Danian | Argentina | Gasteroclupea branisai (219933 219934 219935 219936 219937 219939) | |
| Pu2 | Canada (Saskatchewan) | Diplomystus sp. (14764) | |
| Danian | Bolivia (Mizque) | Gasteroclupea branisai (13501) | |
| Ypresian | Italy | Eoellimmichthys superstes (55239) | |
| Eocene | New Zealand (South Island) | Scombroclupea macropthalma (204226) | |
| Wasatchian | USA (Wyoming) | Diplomystus dentatus (15853) Diplomystus sp. (74615 74616 74617 85026 192130) |