PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Bursa

Gastropoda - Bursidae

Taxonomy

Bursa was named by Roding (1798) [Sepkoski's age data: T Than R Sepkoski's reference number: 705]. It is extant.

It was assigned to Ranellidae by Clark and Arnold (1923); to Cymatiidae by Ladd (1977); to Neotaenioglossa by Sepkoski (2002); and to Bursidae by Oostingh (1935), Mukerjee (1939), MacNeil (1960), Adegoke (1977), Ladd (1982), Perrilliat (1987), Robba et al. (1989), Wilson (1993), Todd (2001), Beu (2005), Beu (2010), Landau and Marques da Silva (2010) and Beu et al. (2012).

Species

B. (Bursa), B. amphitrites, B. asperrima, B. bufonia (syn. B. mammata, B. monitata), B. chipolana, B. chira, B. condita (syn. Tritonium candisatum), B. corrugata (syn. Ranella pustulosa, Ranella caelata, Ranella ponderosa, B. mexicana, Ranella semigranosa), B. cruentata, B. granularis (syn. B. alfredensis, Biplex rubicola, Tritonium jabick, Ranella granifera, B. kowiensis, B. cumingiana, Ranella livida, Ranella affinis, Ranella cubaniana), B. grayana (syn. B. bufoniopsis), B. koperbergae, B. lamarckii (syn. B. angioyorum, B. muehlhaeusseri), B. latitudo, B. natalensis, B. ranelloides (syn. Simpulum papillosum, B. pygmaea, B. finlayi), B. rhodostoma (syn. Ranella paulucciana, Ranella venustula, Ranella thomae, Ranella bergeri), B. rosa (syn. Ranella siphonata), B. rubeta, B. rugosa, B. saundersi, B. scrobilator (syn. Murex nodosus), B. trampasensis, B. victrix, B. washingtoniana

Species lacking formal opinion data

B. scrobiculator

Synonyms

Lampasopsis was named by Jousseaume (1881) [Sepkoski's age data: T Mi R]. It is extant.

It was corrected as Lampadopsis by Fischer (1884); it was reranked as Bursa (Lampadopsis) by Dall (1908); it was synonymized subjectively with Bursa by Beu (2005).

It was assigned to Gastropoda by Fischer (1884); to Bursa by Dall (1908); and to Neotaenioglossa by Sepkoski (2002).
Pseudobursa was named by Rovereto (1899). It is extant.

It was synonymized subjectively with Bursa by Beu (2005).
Annaperenna was named by Iredale (1936). It is extant.

It was synonymized subjectively with Bursa by Beu (2005).
Bufonariella was named by Thiele (1929) [Sepkoski's age data: T Mi R]. It is extant.

It was reranked as Bursa (Bufonariella) by Schmelz (1997); it was synonymized subjectively with Bursa by Beu (2005).

It was assigned to Bursa by Schmelz (1997); and to Neotaenioglossa by Sepkoski (2002).
Colubrellina was named by Fischer (1884) [Sepkoski's age data: T Eo R]. It is extant.

It was reranked as Bursa (Colubrellina) by Ladd (1977) and Perrilliat (1987); it was synonymized subjectively with Bursa by Beu (2005).

It was assigned to Bursa by Ladd (1977) and Perrilliat (1987); and to Neotaenioglossa by Sepkoski (2002).

Synonymy list

Year	Name and author
1798	Bursa Roding
1881	Lampasopsis Jousseaume p. 175
1884	Colubrellina Fischer p. 656
1884	Lampadopsis Fischer p. 656
1899	Pseudobursa Rovereto p. 6
1908	Bursa (Lampadopsis) Dall p. 320
1923	Bursa Clark and Arnold p. 163
1929	Bufonariella Thiele p. 284
1935	Bursa Oostingh p. 61
1936	Annaperenna Iredale p. 310
1939	Bursa Mukerjee p. 54
1960	Bursa MacNeil p. 60
1977	Bursa Adegoke p. 209
1977	Bursa Ladd p. 35
1977	Bursa (Colubrellina) Ladd p. 35
1982	Bursa Ladd p. 41
1987	Bursa Perrilliat p. 15
1987	Bursa (Colubrellina) Perrilliat p. 15
1989	Bursa Robba et al. p. 77
1993	Bursa Wilson p. 226
1997	Bursa (Bufonariella) Schmelz p. 105
2001	Bursa Todd
2002	Bufonariella Sepkoski
2002	Bursa Sepkoski
2002	Colubrellina Sepkoski
2002	Lampasopsis Sepkoski
2005	Bursa Beu p. 18
2010	Bursa Beu p. 41
2010	Bursa Landau and Marques da Silva p. 59
2012	Bursa Beu et al. p. 63

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
phylum	Mollusca
class	Gastropoda

Rank	Name	Author
subclass	Caenogastropoda	(Cox 1959)
—	Sorbeoconcha	(Ponder and Lindberg 1997)
—	Hypsogastropoda	(Ponder and Lindberg 1997)
superorder	Latrogastropoda	Riedel 2000
superfamily	Tonnoidea	(Suter 1913)
family	Bursidae	Thiele 1925
genus	Bursa	Roding 1798

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Bursa Roding 1798

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Subg. Bursa (Bursa) Röding 1798

†Bursa amphitrites Maury 1917

Bursa asperrima Dunker 1862

Bursa bufonia Gmelin 1791

Invalid names: Bursa mammata Röding 1798 [synonym], Bursa monitata Röding 1798 [synonym]

†Bursa chipolana Schmelz 1997

†Bursa chira Olsson 1930

Bursa condita Gmelin 1791

Invalid names: Tritonium candisatum Röding 1798 [synonym]

Bursa corrugata Perry 1811

Invalid names: Bursa mexicana Perrilliat 1963 [synonym], Ranella caelata Broderip 1833 [synonym], Ranella ponderosa Reeve 1844 [synonym], Ranella pustulosa Reeve 1844 [synonym], Ranella semigranosa Lamarck 1822 [synonym]

Bursa cruentata Sowerby 1835

Bursa granularis Röding 1798

Invalid names: Biplex rubicola Perry 1811 [synonym], Bursa alfredensis Turton 1932 [synonym], Bursa cumingiana Dunker 1862 [synonym], Bursa kowiensis Turton 1932 [synonym], Ranella affinis Broderip 1833 [synonym], Ranella cubaniana d'Orbigny 1841 [synonym], Ranella granifera Röding 1798 [synonym], Ranella livida Reeve 1844 [synonym], Tritonium jabick Röding 1798 [synonym]

Bursa grayana Dunker 1862

Invalid names: Bursa bufoniopsis Maury 1917 [synonym]

Bursa koperbergae Van Regteren Altena 1942

Bursa lamarckii Deshayes 1832

Invalid names: Bursa angioyorum Parth 1990 [synonym], Bursa muehlhaeusseri Parth 1990 [synonym]

Bursa latitudo Garrard 1961

Bursa natalensis Coelho and Matthews 1970

Bursa ranelloides Reeve 1844

Invalid names: Bursa finlayi McGinty 1962 [synonym], Bursa pygmaea Kosuge 1979 [synonym], Simpulum papillosum Adams 1870 [synonym]

Bursa rhodostoma Sowerby II 1835

Invalid names: Ranella bergeri Tapparone-Canefri 1881 [synonym], Ranella paulucciana Tapparone-Canefri 1876 [synonym], Ranella thomae d'Orbigny 1841 [synonym], Ranella venustula Reeve 1844 [synonym]

Bursa rosa Perry 1811

Invalid names: Ranella siphonata Reeve 1844 [synonym]

†Bursa rubeta Röding 1798

Bursa rugosa Sowerby II 1835

†Bursa saundersi Adegoke 1977

Bursa scrobilator Linnaeus 1758

Invalid names: Murex nodosus Borson 1825 [synonym]

Bursa trampasensis Clark 1915

†Bursa victrix Dall 1916

†Bursa washingtoniana Weaver 1912

Invalid names: Annaperenna Iredale 1936 [synonym], Bufonariella Thiele 1929 [synonym], Colubrellina Fischer 1884 [synonym], Lampasopsis Jousseaume 1881 [synonym], Pseudobursa Rovereto 1899 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

aragonitec

Locomotion:

actively mobilef

Life habit:

epifaunalf

Diet:

carnivoref

Vision:

limitedc

Created:

2009-09-10 07:59:04

Modified:

2009-09-10 09:59:04

Source: f = family, c = class

References: Hendy et al. 2009, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Selandian to the top of the Holocene or 61.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 59.2 Ma

Collections (182 total)

Time interval	Ma	Country or state	Original ID and collection number
Selandian	61.6 - 59.2	Nigeria	B. saundersi (7791)
Bartonian	41.3 - 38.0	Peru (Piura)	B. chira (52526)
Bartonian	41.3 - 38.0	USA (Oregon)	Cymatium washingtoniana (200858 200863)
Bartonian	41.3 - 38.0	USA (Washington)	B. cowlitzensis, B. washingtoniana (139020) B. washingtoniana (207129) Ranella washingtoniana (52205 52206)
Priabonian	38.0 - 33.9	Peru	B. chira (179594)
Priabonian	38.0 - 33.9	Peru (Ica)	B. chira (100331)
Priabonian - Rupelian	38.0 - 28.1	Peru	B. chira (219109)
Late/Upper Eocene	37.2 - 33.9	Peru	B. sp. (41616)
Late/Upper Eocene	37.2 - 33.9	USA (California)	B. washingtoniana (193039)
Early/Lower Oligocene	33.9 - 28.4	Peru	B. sp. (41619)
Early/Lower Oligocene	33.9 - 28.4	USA (Georgia)	B. vitrix (60835)
Early/Lower Oligocene	33.9 - 28.4	Peru (Piura)	B. sp. (55433)
Late/Upper Oligocene	28.4 - 23.03	Peru (Piura)	B. sp. (64559)
Aquitanian	23.03 - 20.44	France	B. tuberosa (74861)
Early/Lower Miocene	23.03 - 15.97	Grenada	B. rugosa (111216)
Early/Lower Miocene	23.03 - 15.97	India	Argobuccinum affinis (72033) B. elegans (72084 76573 76574 78641)
Early/Lower Miocene	23.03 - 15.97	Venezuela (Falcon)	B. rhodostoma, B. rugosa (41832) B. rugosa (132319)
Burdigalian	20.44 - 15.97	USA (Florida)	B. chipolana (89574 89575 92544 92569) B. chipolana, B. rhodostoma (35018 89576)
Burdigalian	20.44 - 15.97	Venezuela (Falcon)	B. caelata (60144 60705 60710)
Burdigalian	20.44 - 15.97	Venezuela (Falcón)	B. rhodostoma, B. rugosa (96434)
Langhian	15.97 - 13.82	Colombia (La Guajira)	B. rugosa (136885 157830 157858 160970)
Langhian	15.97 - 13.82	Venezuela (Falcon)	B. amphitrites (136469)
Middle Miocene	15.97 - 11.608	Mexico (Michoacan)	B. sp. (66163)
Middle Miocene	15.97 - 11.608	Panama	B. caelata (52173)
Middle Miocene	15.97 - 11.608	Grenada	B. rugosa (58667 58676 58681) B. sp. A (58684)
Middle Miocene	15.97 - 11.608	Trinidad and Tobago	B. ranelloides (136539) B. rhodostoma (96447)
Serravallian	13.82 - 11.62	USA (Florida)	B. amphritrites (111204)
Serravallian - Tortonian	13.82 - 7.246	Italy	B. ranelloides (191624)
Badenian	13.65 - 12.7	Austria	B. sp. (74656)
Badenian	13.65 - 12.7	Poland (Holy Cross Mountains)	B. papillosa, B. nodosa (38823)
Tortonian	11.62 - 7.246	Panama (Darien)	B. rugosa (136122) B. sp. (136007)
Tortonian	11.62 - 7.246	Venezuela	B. rugosa (96470 130974 130976 130977 131306)
Tortonian	11.62 - 7.246	Panama	B. caelata (82116 82128 93176) B. rugosa (92597 92598 136005 136313 145522 145524) B. rugosa, B. amphitrites (92444 92445) B. sp. (82121)
Tortonian	11.62 - 7.246	Dominican Republic	B. amphitrites (58817) B. amphritites (93686)
Tortonian	11.62 - 7.246	Panama (Colon)	B. caelata (145254 145278) B. rugosa (79780 79783 79784 79790 79996 136014 136598)
Late/Upper Miocene	11.608 - 5.333	Ecuador	B. rugosa (131016)
Late/Upper Miocene	11.608 - 5.333	Colombia	B. rugosa (96501)
Late/Upper Miocene	11.608 - 5.333	Ecuador (Esmeraldas)	B. rugosa (136317)
Late/Upper Miocene	11.608 - 5.333	Trinidad and Tobago	B. thomae (60637)
Messinian	7.246 - 5.333	Panama (Bocas)	B. rugosa (135944)
Messinian	7.246 - 5.333	Panama (Bocas del Toro)	B. rugosa (135816 135822)
Messinian	7.246 - 5.333	Dominican Republic	B. amphitrites (75041 89557 89559 89564) B. grayana (84952 84965 89540 93709 93716 94000 131567 131571 131645 131663)
Messinian	7.246 - 5.333	Japan	B. ranelloides (161785)
Messinian	7.246 - 5.333	Panama (Colon)	B. sp. (79855)
Messinian	7.246 - 5.333	Spain	B. sp. (56687)
Messinian - Zanclean	7.246 - 3.6	Dominican Republic	B. amphitrites (89542) B. amphitrites, B. grayana (75038) B. grayana (74967 75036 75037 75039 75040 75043 89561 89562)
Early/Lower Pliocene	5.333 - 3.6	Ecuador (Manabi)	B. sp. (60772)
Early/Lower Pliocene	5.333 - 3.6	Panama (Bocas)	B. rugosa (135839)
Early/Lower Pliocene	5.333 - 3.6	Venezuela	B. caelata (96486) B. rugosa (131315 131317 131322 131323 136387)
Early/Lower Pliocene	5.333 - 3.6	Ecuador (Esmeraldas)	B. rugosa (111211)
Early/Lower Pliocene	5.333 - 3.6	Colombia (Bolívar)	B. scrobilator (136630)
Early/Lower Pliocene	5.333 - 3.6	Dominican Republic	B. amphitrites (58873) B. bufoniopsis (58819 58821)
Early/Lower Pliocene	5.333 - 3.6	Ecuador	B. sp. (131293)
Zanclean	5.333 - 3.6	Italy (Asti)	B. nodosa (86305)
Zanclean	5.333 - 3.6	Dominican Republic	B. amphitrites, B. grayana (74968 80136) B. grayana (93737 93852)
Pliocene	5.333 - 2.588	Fiji	B. nobilis (78073)
Pliocene	5.333 - 2.588	Pakistan	B. sp. (55399)
Pliocene	5.333 - 2.588	Papua New Guinea	B. interrupta, B. tuberculata (78263)
Late/Upper Pliocene	3.6 - 2.588	Venezuela	B. rugosa (79710 111010 111011)
Late/Upper Pliocene	3.6 - 2.588	Jamaica	B. sp. (69093)
Late/Upper Pliocene	3.6 - 2.588	Mexico (Veracruz)	B. corrguata, B. rugosa (96440) B. corrugata (92449) B. corrugata, B. rugosa (92779)
Late/Upper Pliocene	3.6 - 2.588	USA (Florida)	B. amphitrites (79958)
Late/Upper Pliocene	3.6 - 2.588	Costa Rica	B. natalensis, B. rhodostoma, B. rugosa, B. scrobilator (96519) B. scrobilator (92575)
Piacenzian	3.6 - 2.588	Panama (Bocas del Toro)	B. grayana (136522) B. ranelloides (136540)
Piacenzian	3.6 - 2.588	Spain	B. scrobilator (80562)
Piacenzian	3.6 - 2.588	Panama (Bocas)	B. sp. (79999)
Piacenzian	3.6 - 2.588	Italy (Emilia-Romagna)	B. nodosa (178117)
Gelasian	2.588 - 1.806	Italy	B. nodosa (178457)
Gelasian	2.588 - 1.806	Japan (Okinawa)	B. corrugata (42178 42180)
Early/Lower Pleistocene	2.588 - 0.781	Indonesia	B. koperbergae (67971)
Early/Lower Pleistocene	2.588 - 0.781	Panama	B. rugosa (111213 130892)
Early/Lower Pleistocene	2.588 - 0.781	New Zealand (Kermadec Islands)	B. granularis (85086)
Early/Lower Pleistocene	2.588 - 0.781	Spain	B. nodosa (128383)
Pleistocene	2.588 - 0.0117	Costa Rica	B. caelata (135044) B. rugosa (136542)
Pleistocene	2.588 - 0.0117	Venezuela	B. rhodostoma (136541)
Pleistocene	2.588 - 0.0117	Sudan	Ranella affinis (101414)
Pleistocene	2.588 - 0.0117	Ecuador (Manabi)	B. corrugata (136455)
Pleistocene	2.588 - 0.0117	Egypt	Ranella affinis (101633)
Middle Pleistocene	0.774 - 0.129	Spain	B. pustulosa (128324) B. scrobiculator (128384)
Middle Pleistocene	0.774 - 0.129	Vanuatu	B. rubeta, B. bufonia (78043)
Middle Pleistocene	0.774 - 0.129	Angola	B. scrobilator (152645)
Middle Pleistocene	0.774 - 0.129	USA (California)	B. califomica (96894)
Middle Pleistocene - Late/Upper Pleistocene	0.774 - 0.0117	USA (Hawaii)	B. corrugata, B. bufonia (89294)
Late/Upper Pleistocene	0.129 - 0.0117	Portugal (Azores)	B. scrobiculator (89618)
Late/Upper Pleistocene (interglacial)	0.129 - 0.0117	Bahamas	B. sp. (80677)
Late/Upper Pleistocene (interglacial)	0.129 - 0.0117	Jamaica	B. cubaniana (80407)
Late/Upper Pleistocene	0.129 - 0.0117	Spain	B. scrobiculator (127359 128385)
Late/Upper Pleistocene	0.129 - 0.0117	Australia (Western Australia)	B. granularis (63356)
Late/Upper Pleistocene	0.129 - 0.0117	Japan	B. nobilis, B. ranelloides (88886)
Holocene	0.0117 - 0.0	Venezuela	B. corrugata (136357)