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Dacentrurus armatus
Taxonomy
Omosaurus armatus was named by Marsh (1877). Its type specimen is BMNH 46013, a set of postcrania, and it is a 3D body fossil. Its type locality is Swindon Brick and Tile Company brick pit, which is in a Kimmeridgian marine claystone in the Kimmeridge Clay Formation of the United Kingdom. It is the type species of Dacentrurus.
It was recombined as Stegosaurus armatus by Dollo (1889), Gadow (1901); it was considered a nomen dubium by Maidment (2010); it was recombined as Dacentrurus armatus by Lucas (1902), Hennig (1915), Delair (1960), Delair (1973), Thulborn (1975), Galton (1982), Galton (1983), Galton (1985), Galton (1990), Galton (1991), Olshevsky and Ford (1993), Galton (1994), Casanovas-Cladellas et al. (1995), Casanovas-Cladellas et al. (1995), Sanz (2000), Pereda Suberbiola and Galton (2001), Carpenter et al. (2001), Blows (2001), Ruiz-Omeñaca and Canudo (2003), Pereda Suberbiola et al. (2003), Pereda Suberbiola et al. (2005), Mateus (2006), Escaso et al. (2006), Escaso et al. (2007), Maidment et al. (2008), Suñer et al. (2008), Naish and Martill (2008), García-Ramos et al. (2008), Cobos et al. (2008), Butler et al. (2008), Ortega et al. (2009), Pereda Suberbiola et al. (2009), Company et al. (2010), Mateus and Milàn (2010), Barrett and Maidment (2011), Gaete et al. (2011), Pascual et al. (2012), Cobos and Gascó (2013), Cobos et al. (2014), Ulansky (2014), Ulansky (2014), Saitta (2015), Borinder et al. (2016), Galton and Carpenter (2016), Costa and Mateus (2019), Mocho et al. (2019), Maidment et al. (2020), Breeden et al. (2021), Sánchez-Fenollosa et al. (2024), Sánchez-Fenollosa and Cobos (2025).
It was recombined as Stegosaurus armatus by Dollo (1889), Gadow (1901); it was considered a nomen dubium by Maidment (2010); it was recombined as Dacentrurus armatus by Lucas (1902), Hennig (1915), Delair (1960), Delair (1973), Thulborn (1975), Galton (1982), Galton (1983), Galton (1985), Galton (1990), Galton (1991), Olshevsky and Ford (1993), Galton (1994), Casanovas-Cladellas et al. (1995), Casanovas-Cladellas et al. (1995), Sanz (2000), Pereda Suberbiola and Galton (2001), Carpenter et al. (2001), Blows (2001), Ruiz-Omeñaca and Canudo (2003), Pereda Suberbiola et al. (2003), Pereda Suberbiola et al. (2005), Mateus (2006), Escaso et al. (2006), Escaso et al. (2007), Maidment et al. (2008), Suñer et al. (2008), Naish and Martill (2008), García-Ramos et al. (2008), Cobos et al. (2008), Butler et al. (2008), Ortega et al. (2009), Pereda Suberbiola et al. (2009), Company et al. (2010), Mateus and Milàn (2010), Barrett and Maidment (2011), Gaete et al. (2011), Pascual et al. (2012), Cobos and Gascó (2013), Cobos et al. (2014), Ulansky (2014), Ulansky (2014), Saitta (2015), Borinder et al. (2016), Galton and Carpenter (2016), Costa and Mateus (2019), Mocho et al. (2019), Maidment et al. (2020), Breeden et al. (2021), Sánchez-Fenollosa et al. (2024), Sánchez-Fenollosa and Cobos (2025).
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1875 | Omosaurus armatus Owen p. 45 |
| 1876 | Omosaurus armatus Hudleston p. 543 |
| 1876 | Omosaurus armatus Sauvage p. 439 |
| 1877 | Omosaurus armatus Marsh p. 435 |
| 1877 | Omosaurus armatus Owen p. 95 |
| 1878 | Omosaurus armatus Cope p. 57 |
| 1880 | Omosaurus armatus Sauvage p. 523 |
| 1882 | Omosaurus armatus Quenstedt p. 188 |
| 1887 | Osmosaurus armatus Hulke p. 376 |
| 1887 | Omosaurus armatus Hulke p. 700 |
| 1888 | Omosaurus armatus Lydekker p. 177 |
| 1889 | Stegosaurus armatus Dollo p. 680 |
| 1890 | Omosaurus armatus Woodward and Sherborn p. 253 |
| 1890 | Omosaurus armatus Zittel p. 748 |
| 1893 | Omosaurus armatus Seeley p. 54 |
| 1893 | Omosaurus armatus Winkler p. 104 |
| 1896 | Omosaurus armatus Depéret p. 185 |
| 1896 | Omosaurus armatus Mansel-Pleydell p. 119 |
| 1896 | Omosaurus armatus Marsh p. 196 |
| 1901 | Stegosaurus armatus Gadow p. 425 |
| 1901 | Omosaurus armatus Nopcsa p. 214 |
| 1902 | Dacentrurus armatus Lucas p. 435 |
| 1902 | Omosaurus armatus Sauvage p. 408 |
| 1910 | Omosaurus armatus Lull p. 377 |
| 1911 | Omosaurus armatus Nopcsa p. 23 |
| 1915 | Dacentrurus armatus Hennig p. 11 |
| 1924 | Omosaurus armatus Lull p. 242 |
| 1939 | Omosaurus armatus Kuhn p. 113 |
| 1951 | Omosaurus armatus Lapparent and Zbyszewski p. 1127 |
| 1957 | Omosaurus armatus Lapparent and Zbyszewski p. 48 |
| 1960 | Dacentrurus armatus Delair p. 87 |
| 1970 | Omosaurus armatus Swinton p. 241 |
| 1973 | Dacentrurus armatus Delair p. 5 |
| 1975 | Dacentrurus armatus Thulborn p. 89 |
| 1982 | Dacentrurus armatus Galton p. 145 |
| 1983 | Dacentrurus armatus Galton p. 11 |
| 1985 | Dacentrurus armatus Galton p. 213 figs. 2-14 |
| 1990 | Dacentrurus armatus Galton |
| 1991 | Dacentrurus armatus Galton pp. 317-318 |
| 1993 | Dacentrurus armatus Olshevsky and Ford |
| 1994 | Dacentrurus armatus Galton p. 257 |
| 1995 | Dacentrurus armatus Casanovas-Cladellas et al. p. 269 |
| 2000 | Dacentrurus armatus Sanz p. 292 |
| 2001 | Dacentrurus armatus Blows pp. 138-139 |
| 2001 | Dacentrurus armatus Carpenter et al. p. 73 |
| 2001 | Dacentrurus armatus Pereda Suberbiola and Galton p. 150 |
| 2003 | Dacentrurus armatus Pereda Suberbiola et al. p. 144 |
| 2003 | Dacentrurus armatus Ruiz-Omeñaca and Canudo p. 282 |
| 2005 | Dacentrurus armatus Pereda Suberbiola et al. p. 176 |
| 2006 | Dacentrurus armatus Escaso et al. p. 79 |
| 2006 | Dacentrurus armatus Mateus p. 231 |
| 2007 | Dacentrurus armatus Escaso et al. p. 158 |
| 2008 | Dacentrurus armatus Butler et al. p. 7 |
| 2008 | Dacentrurus armatus Cobos et al. p. 90 |
| 2008 | Dacentrurus armatus García-Ramos et al. p. 33 |
| 2008 | Dacentrurus armatus Maidment et al. p. 6 |
| 2008 | Dacentrurus armatus Naish and Martill p. 616 |
| 2008 | Dacentrurus armatus Suñer et al. p. 400 |
| 2009 | Miragaia longicollum Araújo et al. p. 3 fig. 2 |
| 2009 | Miragaia longicollum Mateus p. 144A |
| 2009 | Miragaia longicollum Mateus et al. |
| 2009 | Dacentrurus armatus Ortega et al. p. 48 |
| 2009 | Miragaia longicollum Ortega et al. p. 49 |
| 2009 | Dacentrurus armatus Pereda Suberbiola et al. p. 13 |
| 2010 | Dacentrurus armatus Company et al. p. 248 |
| 2010 | Miragaia longicollum Galton p. 190 |
| 2010 | Dacentrurus armatus Mateus and Milàn p. 253 |
| 2011 | Dacentrurus armatus Barrett and Maidment p. 404 |
| 2011 | Dacentrurus armatus Gaete et al. p. 179 |
| 2012 | Dacentrurus armatus Pascual et al. p. 306 |
| 2013 | Dacentrurus armatus Cobos and Gascó p. 18 |
| 2013 | Miragaia longicollum Ruiz-Omeñaca et al. p. 37 |
| 2014 | Dacentrurus armatus Cobos et al. p. 48 |
| 2014 | Dacentrurus armatus Ulansky p. 7 |
| 2014 | Miragaia longicollum Ulansky p. 7 |
| 2014 | Dacentrurus armatus Ulansky p. 13 |
| 2014 | Miragaia longicollum Ulansky p. 16 |
| 2015 | Dacentrurus armatus Saitta p. 2 |
| 2016 | Dacentrurus armatus Borinder et al. p. 64 |
| 2016 | Dacentrurus armatus Galton and Carpenter p. 200 |
| 2019 | Dacentrurus armatus Costa and Mateus p. 19 |
| 2019 | Miragaia longicollum Costa and Mateus p. 20 |
| 2019 | Dacentrurus armatus Mocho et al. p. 2 |
| 2020 | Dacentrurus armatus Maidment et al. p. 91 |
| 2020 | Miragaia longicollum Maidment et al. p. 92 |
| 2021 | Dacentrurus armatus Breeden et al. p. 21 |
| 2023 | Miragaia longicollum Manitkoon et al. p. 3 |
| 2024 | Miragaia longicollum Burigo and Mateus p. 4 |
| 2024 | Dacentrurus armatus Sánchez-Fenollosa et al. p. 2 |
| 2025 | Dacentrurus armatus Sánchez-Fenollosa and Cobos p. 170 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Dacentrurus armatus Marsh 1877
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Invalid names: Miragaia longicollum Mateus et al. 2009 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| P. M. Galton 1991 | Twelve cervical vertebrae, centra of posterior third of neck and of the dorsal series are massive so the maximum transverse width is greater than the maximum length; in dorsal vertebrae from posterior two-thirds of series, the solid pedicel-like region is short and the minimum angle between each transverse process and the neural splne is 55°. Sacrum consists of seven coossified centra with two dorsosacral vertebrae anteriorly. Centra of anterior caudal vertebrae are massive and the short neural splnes have massive rounded tops. Anterior process of ilium is short and broadens anteriorly; distal part of ischium tapers and is straight in lateral view. Ratio of maximum lengths of humerus to radius is 1 : 0.69, of humerus to ulna is 1 : 0.79, of femur to humerus is 1 : 0.68, and of femur to ilium is 1: 0.85. Dermal armor includes at least small plates with a thick base, two pair of stocky spines with an expanded base, and four pairs of long caudal spines with a small base (modified from GALTON, 1985). | |
| S. C. R. Maidment et al. 2008 | Differs from all other stegosaurs in possessing dorsal vertebral centra that are wider transversely than they are long anteroposteriorly. The dorsal surface ofthe distal ischial shaft is straight, | |
| O. Mateus et al. 2009 (Miragaia longicollum) | Differs from other stegosaurs in the presence of the following autapomorphies: (i) anterior tip of the premaxilla is drawn into a point, (ii) anterolateral rim of the premaxilla projects ventrally, (iii) at least 17 cervical vertebrae, (iv) mid-cervical neural spines possess a notch at their base with an anterior projection dorsal to it, (v) mid and posterior cervical and anterior dorsal neural spines with transversely expanded apices, and (vi) paired, slightly outwardly convex, triangular cervical dermal plates with a notch and projection on the anterodorsal margin.
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| F. Costa and O. Mateus 2019 | Suggested revised diagnosis, modified from [9] with additional characters named herein. Differs from all other stegosaurs in possessing: (i) straight dorsal surface of the distal ischial shaft; (ii) anterior end of the prepubis expanded ventrally; (iii) apex of cervical neural spines expanded posteriorly; (iv) cervical transverse processes borne at mid height of the prezygapophyses. | |
| F. Costa and O. Mateus 2019 (Miragaia longicollum) | Modified from [22]: Differs from other stegosaurs in the presence of the following autapomorphies: (i) anterior tip of the premaxilla is drawn into a point; (ii) anterolateral rim of the premaxilla projects ventrally; (iii) at least 17 cervical vertebrae; (iv) spinopostzygapophyseal lamina extending anteriorly from the epipophyses, accompanied medially by a pair of lower parallel ridges, passing laterally on the neural spine and culminating on an anterior projection on the base of the neural spine with a notch ventral to it (revised autapomorphy from [22]); (v) mid and posterior cervical and anterior dorsal neural spines with trans- versely expanded apices; (vi) paired, slightly inwardly convex, triangular cervical dermal plates with a notch and projection on the posterodorsal margin (revised autapomorphy from [22]); (vii) cervical neural spines are positioned over the anterior half of the centrum and become progressively more anteriorly positioned passing posteriorly on the cervical series; (viii) cervical transverse processes more than half the axial length of the centrum in all but the anteriormost cervical vertebrae; (ix) outline in lateral view of cervical prezygapophyses round posteriorly and straight anteriorly with an anterodorsal notch; (x) closed proximodorsal canal on the ribs of the first caudal vertebra; (xi) progressively more posteriorly inclined neural spines of anterior caudal vertebrae, inclined at less than 45° to the horizontal between Cd8 and Cd11; (xii) neural spine reduced to one fifth the height and width from the 10th to the 12th caudal vertebra, vestigial further posteriorly in the vertebral series; (xiii) presence of longitudinal cord-like ridges in the femur shaft, two posteriorly and one anterolaterally positioned, with distal bifurcation. | |
| S. Sánchez-Fenollosa and A. Cobos 2025 | D. armatus possesses the following autapomorphies (modified from Sánchez-Fenollosa et al. 2025): A premaxilla with (1) an anterior tip that drawn into a point; and (2) an anterolateral margin ventrally projected; a supraoccipital with (3) a posteroventrally orientation with an angle greater than 90° with the dorsal plane of the skull roof (new); a cervical series with (4) at least 17 cervical vertebrae; and (5) at least anterior and mid cervical ribs fused to the vertebrae; cervical vertebrae with (6) two spinopostzygapophyseal laminae that extend anterolaterally from the top of the postzygapophyses to both sides of the base of the neural spine and culminate on its anterior margin; mid and posterior cervical vertebrae with (7) neural spines positioned in the anterior half of the centrum; anterior caudal vertebrae with (8) short neural spines and expanded and rounded apices; an ilium with (9) a wide and short preacetabular process; and (10) a broad base of
the preacetabular process and a smooth curvature between the anterior margin of the sacral yoke and the dorsal margin of the preacetabular process; and a pubis with (11) a dorsoventrally expanded anterior end of the prepubis. |