PBDB Taxon

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Modiolus

Bivalvia - Mytilida - Mytilidae

Taxonomy

Modiola was named by Lamarck (1801) [Sepkoski's age data: D Fame R]. It is extant.

It was reranked as Mytilus (Modiolus) by Forbes (1846); it was corrected as Modiolum by Görges (1941); it was corrected as Modiolus by Glenn (1904), Clark and Arnold (1923), Nagao (1928), Oostingh (1935), Adegoke (1977), Vokes (1980), Sey and Kalacheva (1980), Moore (1983), Moore (1984), Ward and Blackwelder (1987), Squires and Goedert (1991), Lamprell and Healy (1998), Todd (2001), Sepkoski (2002), Rosenberg (2005), El Qot (2006), Mikkelsen (2006), Scholz et al. (2008), Marquez-Aliaga et al. (2010) and Ros-Franch et al. (2014).

It was assigned to Mytilus by Forbes (1846); to Mytiloida by Sepkoski (2002); to Modiolinae by Adegoke (1977), Moore (1983), Squires and Goedert (1991) and Rosenberg (2005); and to Mytilidae by Lamarck (1799), Stanton (1895), Glenn (1904), Clark and Arnold (1923), Nagao (1928), Oostingh (1935), Görges (1941), Vokes (1980), Sey and Kalacheva (1980), Moore (1984), Ward and Blackwelder (1987), Lamprell and Healy (1998), Todd (2001), El Qot (2006), Mikkelsen (2006), Scholz et al. (2008), Marquez-Aliaga et al. (2010) and Ros-Franch et al. (2014).

Species

Modiola (Lithophagus), Modiola (Septiola), Modiola amygdalina, Modiola angustissima, Modiola barbata, Modiola bolodekensis, Modiola costulata, Modiola decliva, Modiola exbrocchii, Modiola flemischi, Modiola fornicata, Modiola goldfussi, Modiola gracilior, Modiola hartleyensis, Modiola houstonia, Modiola klipsteinii, Modiola lidarensis, Modiola longaticensis, Modiola marmorata, Modiola minuta, Modiola myoconchaeformis, Modiola nevadensis, Modiola ocrophaius, Modiola pantanellii, Modiola pedernalis, Modiola pictinatus, Modiola potomacensis, Modiola praerupta, Modiola pueblensis, Modiola raibliana, Modiola ravni, Modiola rhaetica, Modiola rhombea, Modiola sanharica, Modiola schafhaeutli, Modiola siliqua, Modiola solenoides, Modiola subcarinatus, Modiola sulcatus, Modiola texana, Modiola triquaetraeformis, Modiola tulipea, Modiola visgnolae, Modiola vomer, M. (Cyranus), M. (Gibbomodiola), M. (Inoperna), M. (Modiolus), M. (Modiolusia), M. (Strimodiolus), M. addicotti, M. adelaidensis, M. aequalis, M. alveolana, M. ameghinoi, M. americana, M. americanus, M. anatinus (syn. Modiola cuneata), M. archiskanni, M. areolatus, M. auriculatus, M. ausiensis, M. balongensis, M. bipartitus, M. buckmani, M. capax, M. carpenteri (syn. M. fornicata, Volsella fornicata), M. cawcawensis, M. columbianus, M. dalli, M. dimidiatus, M. ducatelii, M. ducatellii (syn. Modiola inflata), M. ellipticus, M. eugenensis, M. extensus, M. friesenbichlerae, M. frugi, M. gejiuensis, M. gerthi, M. gibbus, M. giganteus, M. gigantoides, M. gigantus, M. gracilis, M. grammatus, M. guiyangensis, M. hollicki, M. ionensis, M. jaworskii, M. kelsoensis, M. koneckii, M. kutinskensis, M. latecaudatus, M. ligeriensis, M. magnus, M. major, M. meeki, M. micropteris, M. minimus, M. muensteri, M. omanensis, M. ornatus, M. paronai, M. philippinarum, M. pliciferus, M. porterensis, M. pulchellus, M. pygmaeus (syn. Mytilus subpygmaeus, Mytilus pygmaeus), M. raubensis, M. restorationensis, M. reversus, M. salzstettensis, M. sandbergeri, M. semilaevigatus, M. silentiensis, M. solea, M. sookensis, M. speciosus, M. stantoni, M. strajeskianus, M. subsimplex, M. tenuistriatus, M. trigonalis, M. trinominata (syn. M. inflatus), M. triquetra, M. typicus, M. virginicus, M. wajampolkensis, M. weiyuanensis, M. willapaensis, M. yunnanensis, Mytilus (Modiolus) cypris, Mytilus (Modiolus) pulcher

Synonymy list

Year	Name and author
1799	Modiola Lamarck
1801	Modiola Lamarck
1846	Mytilus (Modiolus) Forbes p. 152
1895	Modiola Stanton p. 48
1904	Modiolus Glenn p. 365
1923	Modiolus Clark and Arnold p. 143
1928	Modiolus Nagao p. 45
1935	Modiolus Oostingh p. 161
1941	Modiolum Görges p. 159
1977	Modiolus Adegoke p. 229
1980	Modiolus Sey and Kalacheva
1980	Modiolus Vokes p. 26
1983	Modiolus Moore p. A74
1984	Modiolus Moore p. 26
1987	Modiolus Ward and Blackwelder p. 129
1991	Modiolus Squires and Goedert p. 413
1998	Modiolus Lamprell and Healy p. 80
2001	Modiolus Todd
2002	Modiolus Sepkoski
2005	Modiolus Rosenberg
2006	Modiolus El Qot p. 20
2006	Modiolus Mikkelsen
2008	Modiolus Scholz et al.
2010	Modiolus Marquez-Aliaga et al. p. 193
2014	Modiolus Ros-Franch et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
phylum	Mollusca
class	Bivalvia

Rank	Name	Author
—	Eubivalvia
subclass	Autobranchia	(Groblen 1894)
infraclass	Pteriomorphia	(Beurlen 1944)
—	Mytilomorphi	Ferussac 1822
order	Mytilida	(Rafinesque 1815)
superfamily	Mytiloidea	Rafinesque 1815
family	Mytilidae	Rafinesque 1815
genus	Modiolus	(Lamarck 1801)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Modiolus Lamarck 1801

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Subg. Modiola (Lithophagus) Megerle 1811

†Lithophagus alpinus Zittel 1866

†Lithophagus oviformis Gabb 1864

Lithophagus plumula Hanley 1844

†Lithophagus priscus Giebel 1856

†Lithophagus triadicus Bittner 1901

Subg. †Modiola (Septiola) Bittner 1895

†Modiola (Septiola) subcarinata Bittner 1895

†Modiola amygdalina Phillips 1841

†Modiola angustissima Newton 1895