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Auroraceratops rugosus

Osteichthyes

Taxonomy
Auroraceratops rugosus was named by You et al. (2005). Its type specimen is IG-2004-VD-001, a skull, and it is not a trace fossil. Its type locality is Auroraceratops type, Gongpoquan Basin, which is in an Albian fluvial-lacustrine horizon in the Zhonggou Formation of China.

Synonymy list
YearName and author
2005Auroraceratops rugosus You et al.
2007Auroraceratops rugosus You et al. p. 898
2009Auroraceratops rugosus Jin et al. p. 204
2009Auroraceratops rugosus Tanoue et al. p. 433
2010Auroraceratops rugosus Amiot et al. p. 355
2010Auroraceratops rugosus You et al. p. 66
2012Auroraceratops rugosus Tanoue et al. p. 125
2012Auroraceratops rugosus You et al. p. 171
2014Auroraceratops rugosus Hedrick et al. p. 322
2014Auroraceratops rugosus Tanoue and Okazaki p. 80
2015Auroraceratops rugosus Han et al. p. 1
2019Auroraceratops rugosus Li et al. p. 70
2019Auroraceratops rugosus Morschhauser et al. p. 37–38
2019Auroraceratops rugosus Morschhauser et al. p. 78
2019Auroraceratops rugosus Suarez et al. p. 22
2019Auroraceratops rugosus You et al. p. 1
2020Auroraceratops rugosus Yu et al. p. 1
2022Auroraceratops rugosus Son et al. p. 7

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
RankNameAuthor
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Ornithischia()
Genasauria
Cerapoda
Marginocephalia
Ceratopsia()
infraorderNeoceratopsia
genusAuroraceratops
speciesrugosus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Auroraceratops rugosus You et al. 2005
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Diagnosis
ReferenceDiagnosis
E. M. Morschhauser et al. 2019A non-coronosaur neoceratopsian diagnosed by the following autapomorphies: inflated premaxillary teeth; a fungiform expansion of the lacrimal; large tuber caudodorsally on the dentary near the contact with the surangular; and tubercle on the lateral face of the dentary about the mid- dle of the mandible. This last character, a low, broad-based protrusion, is in contrast to the narrow-based, craniocaudally longer and dorsoventrally taller dentary flange seen in Psittacosaurus (Sereno, 2010), the unusual curving structure seen in Chaoyangsaurus (Tanoue et al., 2010), and the structure seen in Liaoceratops (Xu et al., 2006). The fungiform expansion of the dorsal end of the lacrimal is present in many basal neoceratopsians; however, only Auroraceratops has this expansion accentuated by the ventral suture of the lacrimal traveling parallel to and just ventral to the dorsal margin of the antorbital fossa. Auroraceratops is also diagnosed by the following unique combination of characters: rugosity on the dentary, jugal, surangular, and sometimes the postorbital (a derived feature seen in chaoyangsaurids); surangular lacking a continuous ridge (character plesiomorphic to Neoceratopsia, shared only with Liaoceratops within Neoceratopsia); a quadrate with a large neurovascular foramen near the articulation with the quadratojugal (apomorphic, shared with Liaoceratops); larger caudal tuber on the surangular (shared with Archaeoceratops); broad nasals (plesiomorphic for Neoceratopsia, shared with Yinlong, Psittacosaurus, and Stegoceras); and elongate caudal process of the pterygoid
obscuring the basipterygoid-basisphenoid suture from ventral view (derived, shared with Leptoceratops and ZPAL MgD-I/156).
E. M. Morschhauser et al. 2019A non-coronosaur basal neoceratopsian diagnosed by the following autapomorphies: inflated premaxillary teeth; a fungiform expansion of the lacrimal; large tuber caudodorsally on the dentary near the contact with the surangular; and tubercle on the lateral face of the dentary about the middle of the mandible. This last character is a low, broad-based protrusion, in contrast to the narrow-based, craniocaudally longer and taller dentary flange seen in Psittacosaurus (Sereno, 2010), the unusual curving structure seen in Chaoyangsaurus (Tanoue et al., 2010), and the structure seen in Liaoceratops (Xu et al., 2002). The expansion of the dorsal end of the lacrimal is present in many basal neoceratopsians; however, only Auroraceratops has this expansion accentuated by the ventral suture of the lacrimal traveling parallel to and just ventral to the curved dorsal margin of the antorbital fossa. Auroraceratops is also diagnosed by the following unique combination of characters: rugosity on the dentary, jugal, surangular, and sometimes the postorbital (derived relative to Neoceratopsia, independent origin from chaoyangsaurids); surangular lacking a continuous ridge (character plesiomorphic to Neoceratopsia, shared only with Chaoyangsauridae and Liaoceratops within Neoceratopsia); a quadrate with a large neurovascular foramen near the articulation with the quadratojugal (shared with Liaoceratops and ZPAL MgD I/156); larger caudal tuber on the surangular (shared with Archaeoceratops); broad nasals (plesiomorphic for Neoceratopsia, shared with Yinlong, Psittacosaurus, and Stegoceras); and elongate caudal process of the pterygoid obscuring the basipterygoid-basisphenoid suture from ventral view (derived relative to Neoceratopsia, shared with Leptoceratops and ZPAL MgD I/156).
E. M. Morschhauser et al. 2019(Numbers match those in Figs. 1–3.) A non-coronosaur basal neoceratopsian diagnosed by the following autapomorphies: inflated premaxillary teeth (a1); a fungi- form expansion of the lacrimal (a2); large tuber caudodorsally on the dentary near the contact with the surangular (a3); and tubercle on the lateral face of the dentary at the middle of the mandible (a4). This last character is a low, broad-based protrusion and is in contrast to the narrow-based, craniocaudally longer and taller dentary flange seen in Psittacosaurus (Sereno, 2010), the unusual curving structure seen in Chaoyangsaurus (Tanoue et al., 2010), and the structure seen in Liaoceratops (Xu et al., 2006). The fungiform expansion of the dorsal end of the lacrimal is present in many basal neoceratopsians; however, only in Auroraceratops is this expansion accentuated by the ventral suture of the lacrimal lying parallel to and just ventral to the dorsal margin of the antorbital fossa. Auroraceratops is also diagnosed by the following unique combination of characters: rugosity on the dentary, jugal, surangular, and sometimes the postorbital (a derived feature seen in chaoyangsaurids) (s1); surangular lacking a continuous ridge (character plesiomorphic to Neoceratopsia, shared only with Liaoceratops within Neoceratopsia) (s2); a quadrate with a large neurovascular foramen near the articulation with the quadratojugal (shared with Liaoceratops) (s3); larger caudal tuber on the surangular (shared with Archaeoceratops) (s4); broad nasals (ple- siomorphic for Neoceratopsia, shared with Yinlong, Psittacosaurus, and Stegoceras) (s5); and elongate caudal pro- cess of the pterygoid obscuring the basipterygoid-basisphenoid suture from ventral view (derived, shared with Leptoceratops and ZPAL MgD-I/156) (s6).