Chekarda, Sylva River (PIN collection 4987): Kungurian, Russian Federation
collected by V. G. Novokshonov 1989-2000

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Reculida - Geinitziidae
Permoshurabia kungurica n. sp. Aristov and Rasnitsyn 2015
Aristov and Rasnitsyn 2015 1 specimen
PIN 4987/1000 (2 measurements)
Insecta - Reculida - Sylvaphlebiidae
Sylvaphlebia fucata n. sp. Aristov 2004
1 specimen
PIN 4987/12 (1 measurement)
Sylvophenoptera fimbriata n. sp. Aristov 2004
1 specimen
PIN 4987/14
Insecta - Paoliida - Anthracoptilidae
Graticladus apiatus n. sp. Aristov and Rasnitsyn 2015
Aristov and Rasnitsyn 2015 1 specimen
PIN 4987/1002
Insecta - Protorthoptera - Sheimiidae
Parasheimia rotundata n. sp. Aristov 2004
3 specimens
PIN 4987/30-32 (6 measurements)
Parasheimia truncata n. gen., n. sp. Aristov 2004
3 specimens
PIN 4987/27-29 (6 measurements)
Sheimia tshekardensis n. sp. Aristov 2004
2 specimens
PIN 4987/25, 26 (4 measurements)
Insecta - Grylloblattodea - Euremiscidae
Euremisca splendens Zalessky 1951
Aristov 2011 6 specimens
PIN 4987/96, 97, 711, 712, 817, 818
Insecta - Grylloblattodea - Bajanzhargalanidae
Sylvafossor forcipatus n. gen., n. sp. Aristov 2004
1 specimen
PIN 4987/11 (3 measurements)
Insecta - Grylloblattodea
Tshekardites curculiuculus n. sp. Aristov 2004
1 specimen
recombined as Sylvalitoralis curculiuculus
PIN 4987/48 (1 measurement)
Tshekardites calcomessor n. sp. Aristov 2004
1 specimen
recombined as Sylvalitoralis calcomessor
PIN 4987/47 (1 measurement)
Tshekardites lientericus n. sp. Aristov 2004
1 specimen
recombined as Tshebardanympha lientericus
PIN 4987/46 (1 measurement)
Insecta - Grylloblattodea - Liomopteridae
Kungurolioma cancellata n. gen., n. sp. Aristov 2004
2 specimens
PIN 4987/39, 40 (4 measurements)
Khosarophlebia sylvaensis Martynov 1940
2 specimens
PIN 4987/34, 35 (3 measurements)
Parasylvaella minor n. sp. Aristov 2004
1 specimen
PIN 4987/38 (1 measurement)
Parasylvaella umbra n. gen., n. sp. Aristov 2004
1 specimen
PIN 4987/37 (2 measurements)
Uralioma maxima n. sp. Aristov 2004
1 specimen
PIN 4987/45 (1 measurement)
Cerasopterum impurum n. sp. Aristov 2004
1 specimen
recombined as Tapopterum impurum
PIN 4987/44 (1 measurement)
Ornaticosta novokshonovi n. sp. Aristov 2004
1 specimen
PIN 4987/36 (3 measurements)
Insecta - Grylloblattodea - Lemmatophoridae
Culiciforma formosa Aristov 2004
1 specimen
PIN 4987/1
Sylvaprisca focaleata n. sp. Aristov 2004
1 specimen
PIN 4987/2 (3 measurements)
Tshekardites gracilis Aristov 2004
1 specimen
recombined as Lemmatonympha gracilis
PIN 4987/49 (1 measurement)
Artinska infigurabilis n. sp. Aristov 2004
6 specimens
PIN 4987/3-8 (6 measurements)
Insecta - Grylloblattodea
Sylvamicropteron harpax n. gen., n. sp. Aristov 2004
1 specimen
PIN 1987/50 (2 measurements)
Insecta - Cnemidolestodea - Protembiidae
Tshekardomina imbecillissima Aristov 2004
1 specimen
PIN 4987/24
Tshekardomina imbecilla n. sp. Aristov 2004
2 specimens
PIN 4987/22, 23 (6 measurements)
Insecta - Cnemidolestodea - Neraphidiidae
Neraphidia mitis Novokshonov and Novokshonova 1997
1 specimen
PIN 4987/10
Insecta - Cnemidolestodea - Tillyardembiidae
Kungurembia brevicervix Aristov 2004
4 specimens
PIN 4987/15-18
Kungurembia pallida n. sp. Aristov 2004
2 specimens
PIN 4987/19-21 (5 measurements)
Tillyardembia sp. Zalessky 1937
Khramov et al. 2023 1 specimen
PIN 4987/669
Tillyardembia antennaeplana Zalessky 1937
Aristov and Rasnitsyn 2010 7 specimens
PIN 4987/55, 54, 664, 668, 686, 688, 710
Insecta
Curvilarva tshekardensis n. gen., n. sp. Novokshonov 2004
Novokshonov 2004 3 specimens
PIN 4987/109, 110, 111 (1 measurement)
Insecta - Protomeroptera - Protomeropidae
Protomeropidae indet. Tillyard 1926
Khramov et al. 2022 2 specimens
PIN 4987/965, 966
Insecta - Megasecoptera - Protohymenidae
Protohymen novokshonovi n. sp. Pecharová and Prokop 2018
Pecharová and Prokop 2018 1 specimen
PIN 4987/115
Insecta - Engellestidae
Engellestes chekardensis Nel et al. 2012
Felker 2023 1 specimen
PIN 4987/831
Insecta - Eoblattida - Idelinellidae
Euryptilon cuculiophoris Aristov 2002
Aristov and Rasnitsyn 2012 1 specimen
recombined as Strigulla cuculiophora
PIN 4987/821 (3 measurements)
Insecta - Eoblattida - Euryptilonidae
Euryptilon blattoides Martynov 1940
Aristov 2011 4 specimens
PIN 4987/713-716
Insecta - Eoblattida - Mesorthopteridae
Sylviodes perloides Martynov 1940
1 specimen
PIN 4987/33 (1 measurement)
Insecta - Eoblattida - Doubraviidae
Cerasopterum megakhosaroides n. sp. Aristov 2004
3 specimens
recombined as Koshelevka megakhosaroides
PIN 4987/41, 42, 43 (6 measurements)
Insecta - Eoblattida - Atactophlebiidae
Novokshonovus ignoratus n. gen., n. sp. Aristov and Rasnitsyn 2015
Aristov and Rasnitsyn 2015 2 specimens
PIN 4987/811, 810 (1 measurement)
see common names

Geography
Country:Russian Federation State/province:Suksun
Coordinates: 57.0° North, 57.7° East (view map)
Paleocoordinates:30.6° North, 45.4° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Permian Epoch:Cisuralian
Stage:Kungurian 10 m.y. bin:Permian 2
Key time interval:Kungurian
Age range of interval:283.50000 - 273.01000 m.y. ago
Stratigraphy
Formation:Koshelevka
Stratigraphic resolution:group of beds
Stratigraphy comments: Late Kungurian
Lithology and environment
Primary lithology: lithified "siliciclastic"
Environment:deltaic indet. Tectonic setting:foreland basin
Geology comments: Uralian foreland basin. Alluvial to marine delta-fan environment, close to the sea, deposited under arid climate conditions.
Taphonomy
Modes of preservation:mold/impression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Collectors:V. G. Novokshonov Collection dates:1989-2000
Metadata
Also known as:Tshekarda
Database number:122753
Authorizer:M. Clapham Enterer:J. Karr, M. Clapham, L. Edwards
Modifier:M. Clapham Research group:paleoentomology
Created:2012-01-05 20:00:06 Last modified:2013-11-25 13:02:06
Access level:the public Released:2012-01-05 20:00:06
Creative Commons license:CC BY
Reference information

Primary reference:
34932. D. S. Aristov. 2004. The Fauna of Grylloblattid Insects (Grylloblattida) of the Lower Permian Locality of Tshekarda. Paleontological Journal 38:S80-S145 [M. Clapham/J. Karr]

Secondary references:

48771 D. S. Aristov. 2011. Morfologiya nekotorykh Permiskikh Grylloblattida (Insecta) iz mestonakhozhdeniya Chekarda (Permskiy Kray). Sovremennaya Paleontologiya: Klassicheskie i Noveyshie Metody 2011:25-34 [M. Clapham/M. Clapham]
39146 D. S. Aristov and A. P. Rasnitsyn. 2010. The family Tillyardembiidae Zalessky, 1938 and the system of the plecopteroid insects. Russian Entomological Journal 18(4):257-264 [M. Clapham/J. Karr/M. Clapham]
39962 D. S. Aristov and A. P. Rasnitsyn. 2012. Revision of the family Idelinellidae, with a review of the Permian Eoblattida (Insecta). Paleontological Journal 46:49-60 [M. Clapham/M. Clapham]
52003 D. S. Aristov and A. P. Rasnitsyn. 2014. New Eoblattida from the Permian of Russia and the United States and the Origin of earwigs (Insecta: Eoblattida, Forficulida). Paleontological Journal 48:407-413 [M. Clapham/M. Clapham]
55207 D. S. Aristov and A. P. Rasnitsyn. 2015. New insects from the Kungurian of Tshekarda fossil site in Permian territory of Russia. Russian Entomological Journal 24:17-35 [M. Clapham/M. Clapham]
87497 A. S. Felker. 2023. New taxa of damselflies (Protozygoptera) from the Permian of the East European Platform. Paleontological Journal 57:857-914 [M. Clapham/M. Clapham]
84935 A. V. Khramov, T. Foraponova, and P. Węgierek. 2023. The earliest pollen-loaded insects from the Lower Permian of Russia. Biology Letters 19(20220523) [M. Clapham/M. Clapham]
83713 A. V. Khramov, S. V. Naugolnykh, and P. Wegierek. 2022. Possible long-proboscid insect pollinators from the Early Permian of Russia. Current Biology 32:3815-3820 [M. Clapham/M. Clapham]
40007 V. G. Novokshonov. 2004. The First Mecopteroids (Insecta: Papilionidea = Mecopteroidea) and the Origin of Scorpionflies (Panorpida = Mecoptera), with Description of a Legless Eruciform Larva from the Lower Permian of Tshekarda. Paleontological Journal 38(2):204-213 [M. Clapham/L. Edwards]
64642 M. Pecharová and J. Prokop. 2018. The morphology of mouthparts, wings and genitalia of Paleozoic insect families Protohymenidae and Scytohymenidae reveals new details and supposed function. Arthropod Structure & Development 47:117-129 [M. Clapham/M. Clapham]