Sample 3, Titusville Site (0-60 cm): Tarantian, Pennsylvania

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Bembidion morulum LeConte 1863
1 specimen
Insecta - Hemiptera - Saldidae
2 specimens
Insecta - Coleoptera - Scarabaeidae
Aegialia sp. Latreille 1807
2 specimens
Aphodius sp. Hellwig 1798
2 specimens
Insecta - Coleoptera - Helophoridae
Helophorus sp. (Fabricius 1775)
5 specimens
original and current combination Elophorus
Helophorus sempervarians Angus 1970
5 specimens
Insecta - Coleoptera - Hydrophilidae
Hydrophilidae indet. Latreille 1802
4 specimens
Enochrus sp. Thomson 1859
2 specimens
Insecta - Coleoptera - Staphylinidae
Staphylinidae indet. (Latreille 1802)
20 specimens
original and current combination Staphyliniae
Scydmaenidae indet. (Leach 1815)
2 specimens
spelled with current rank as Scydmaeninae
Mycetoporus sp. Mannerheim 1830
1 specimen
Tachinus tachyporoides Horn 1877
1 specimen
recombined as Nitidotachinus tachyporoides
Stenus sp. Latreille 1796
126 specimens
Quedius sp. Stephens 1829
1 specimen
Philonthus sp. Curtis 1829
1 specimen
Omaliinae indet. MacLeay 1825
9 specimens
Microedus austinianus LeConte 1874
1 specimen
Eucnecosum sp. Reitter 1909
5 specimens
Eucnecosum brunnescens (Sahlberg 1871)
1 specimen
Olophrum consimile (Gyllenhal 1810)
6 specimens
Olophrum rotundicolle (Sahlberg 1817)
22 specimens
Olophrum boreale (Paykull 1792)
1 specimen
Pycnoglypta sp. Thomson 1858
3 specimens
Pycnoglypta aptera or Pycnoglypta lurida
Pycnoglypta aptera Campbell 1983
1 specimen
Euaesthetinae indet. Thomson 1859
2 specimens
Bledius sp. Leach 1819
3 specimens
Pselaphidae indet. (Latreille 1802)
1 specimen
Pselaphii
Thanatophilus sp. Leach 1815
1 specimen
Aleocharinae indet. (Fleming 1821)
33 specimens
Gymnusa sp. Gravenhorst 1806
4 specimens
Insecta - Coleoptera - Hydraenidae
Limnebiidae indet.
1 specimen
spelled with current rank as Limnebiini
Ochthebius sp. Leach 1815
1 specimen
Insecta - Coleoptera - Byrrhidae
Cytilus alternatus (Say 1825)
1 specimen
Insecta - Coleoptera - Elateridae
Elateridae indet. Leach 1815
1 specimen
Insecta - Coleoptera - Cantharidae
Podabrus sp. Leconte 1850
1 specimen
Insecta - Coleoptera - Curculionidae
Phloeotribus piceae Swaine 1911
1 specimen
Carphoborus andersoni Swaine 1919
2 specimens
Polygraphus rufipennis (Kirby 1837)
1 specimen
Insecta - Coleoptera - Carabidae
Carabidae indet. (Latreille 1802)
2 specimens
Patrobus sp. Dejean 1821
3 specimens
Patrobus stygicus Chaudoir 1872
3 specimens
Patrobus septentrionis Dejean 1828
1 specimen
Trechus crassiscapus Lindroth 1955
3 specimens
recombined as Trechus (Trechus) crassiscapus
Bembidion informal incertum group
1 specimen
Bembidion mutatum (Gemminger and Harold 1868)
1 specimen
recombined as Bembidion (Bembidion) mutatum
Dyschirius informal integer group
1 specimen
Diacheila arctica (Gyllenhal 1810)
1 specimen
Blethisa catenaria Brown 1944
1 specimen
Elaphrus clairvillei (Kirby 1837)
1 specimen
recombined as Elaphrus (Neoelaphrus) clairvillei
Insecta - Coleoptera - Dytiscidae
Dytiscidae indet. (Leach 1815)
2 specimens
Hydroporus sp. Clairville 1806
2 specimens
Insecta - Hymenoptera - Formicidae
Formicidae indet. (Latreille 1802)
3 specimens
original and current combination Formicariae
see common names

Geography
Country:United States State/province:Pennsylvania
Coordinates: 41.6° North, 79.6° West (view map)
Paleocoordinates:41.6° North, 79.6° West
Basis of coordinate:stated in text
Geographic resolution:hand sample
Time
Period:Quaternary Epoch:Pleistocene
Stage:Late/Upper Pleistocene 10 m.y. bin:Cenozoic 6
Key time interval:Tarantian
Age range of interval:0.12900 - 0.01170 m.y. ago
Stratigraphy
Formation:Titusville Till
Stratigraphy comments: slightly below sample 1 which is dated as 39070 years old
Lithology and environment
Primary lithology: unlithified peat
Environment:mire/swamp Tectonic setting:non-subsiding area
Taphonomy
Modes of preservation:body
Size of fossils:mesofossils
Temporal resolution:time-averaged
Collection methods and comments
Collection methods:selective quarrying,sieve
Reason for describing collection:paleoecologic analysis
Collection method comments: In 1990, 174 kg of peat and silt from six samples was collected from the three organic horizons at depths of 0–60, 120–155, and 245–255 cm, respectively (Fig. 2). The 0-cm data position was defined at the top of the upper organic horizon. Chitinous remains of insects were recovered from the samples using a kerosene flotation technique (Ashworth, 1979). Disaggregation of highly compressed peats was aided by soaking them in a 5% NaOH solution. Arthropod remains were abundant and well preserved in all samples. All specimens, including those examined by Totten and Coope in 1967, were mounted onto micropaleontological slides and reposited in the Quaternary Entomology Laboratory at North Dakota State University.
Metadata
Database number:137205
Authorizer:M. Clapham Enterer:J. Karr
Modifier:M. Clapham Research group:paleoentomology
Created:2012-12-12 09:58:02 Last modified:2015-12-15 18:34:48
Access level:the public Released:2012-12-12 09:58:02
Creative Commons license:CC BY
Reference information

Primary reference:

43873. S. Cong, A. C. Ashworth, D. P. Schwert and S. M. Totten. 1996. Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania. Quaternary Research 45:216-225 [M. Clapham/J. Karr/J. Karr]