Khermeen Tsav I, Red Beds [PMPE]: Middle Campanian - Late Campanian, Mongolia
collected by PMPE 1970–1971

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Hoplocercidae
Gradzinski et al. 1977 1 specimen
ZPAL MgR-III/35 (holotype skull with mandibles)
Reptilia - Priscagamidae
Alifanov 1996 1 individual
ZPAL MgR/III-72 (holotype skull) - this is the same specimen number as a skull referred to priscagama gobiensis by Borsuk-Bialynicka & Moody (1984)
Borsuk-Bialynicka and Moody 1984 5 individuals
ZPAL MgR/III-32 (holotype), 31, 72, 33, 83 (partial skulls)
Reptilia - Polyglyphanodontidae
Sulimski 1975 14 specimens
ZPal MgR-III/7–11, 13, 15–18, 20–21, 28–29
Sulimski 1975 2 specimens
ZPal MgR-III/6
Sulimski 1975 1 specimen
ZPal MgR-III/24
Reptilia
Borsuk-Bialynicka 1988 3 specimens
ZPAL MgR-III/37 (holotype skull), 38 (skull)
Borsuk-Bialynicka 1988 4 specimens
ZPAL MgR-III/40 (holotype skull), 41, 43 (large skulls), 36 (small skull)
Borsuk-Bialynicka 1985 1 specimen
ZPAL Mg-R-III/34 (holotype)
Reptilia - Dinosauria
Gradzinski and Jerzykiewicz 1972
eggs
Reptilia - Elongatoolithidae
Sabath 1991 1 specimen
ZPAL MgO-III/18a-c
Reptilia - Oviraptoridae
Gradzinski and Jerzykiewicz 1972
small theropods (2 measurements)
Osmólska 1976
    = Conchoraptor gracilis Barsbold 1986
Kundrát and Janácek 2007
ZPAL MgD-I/95
Reptilia
Elzanowski 1981 7 specimens
ZPAL MgR-I/33–34, 88–91
Sabath 1991 5 specimens
ZPAL MgO-III/10a-b, 11a-c, 12, 13, 14
Reptilia - Protoceratopsidae
Sabath 1991 1 specimen
ZPAL MgO-III/9
35 individuals
"probably Protoceratops andrewsi" (1 measurement)
Maryanska and Osmólska 1975
ZPAL MgD-I/126
Reptilia - Phaceloolithidae
Mikhailov et al. 1994 1 specimen
ZPAL MgOv-I/16
Mammalia - Multituberculata - Djadochtatheriidae
Kielan-Jaworowska 1974 2 individuals
ZPAL MgM-I/78 (holotype, complete skull), I/79 (damaged cranium)
Hurum and Kielan−Jaworowska 2008
PM 120/107 (partial, associated skeleton)
Kielan-Jaworowska et al. 2003 1 specimen
PM 120/108, partial skull
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 43.3° North, 99.8° East (view map)
Paleocoordinates:37.5° North, 95.5° East (Wright 2013)
Basis of coordinate:stated in text
Altitude:1200 meters
Geographic resolution:small collection
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Campanian 10 m.y. bin: Cretaceous 7
*Period:Late/Upper Cretaceous *Epoch:Senonian
*International age/stage:Middle Campanian
Key time interval: Middle Campanian - Late Campanian
Age range of interval: 83.6 - 72.2 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Baruungoyot
Stratigraphic resolution:group of beds
Stratigraphy comments: = "Khermeen Tsav Formation" = "Red Beds of Khermeen Tsav" = "Protoceratops horizon". Collection from two beds within the formation, close together near middle of section; cannot be separated based on reported sample.
Lithology and environment
Primary lithology:tabular,planar lamination,concretionary poorly lithified silty,calcareous sandstone
Includes fossils?Y
Lithology description: vivid red, poorly cemented sandstone, with sandy concretions of variable size, small, white calcareous concretions occur. Some silt and clay intercalations. Cross-stratification is rare, but horizontally layered and massive beds are common. Common trace fossils - sediment-filled tubes - within the Red Beds.
Environment:eolian indet.
Taphonomy
Modes of preservation:body
Degree of concentration:-multiple events
Size of fossils:macrofossils
Preservation of anatomical detail:good
Articulated whole bodies:none
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Temporal resolution:time-averaged
Spatial resolution:autochthonous
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:selective quarrying,surface (float),surface (in situ),mechanical,field collection
Collection size:35 specimens
Reason for describing collection:taxonomic analysis
Collectors:PMPE Collection dates:1970–1971
Collection method comments: Many other taxa are present, but list includes Khermeen Tsav II localities and they cannot be reliably separated based on this paper.
Metadata
Also known as:Polish-Mongolian Palaeontological Expedition, Khermin Tsav
Database number:13777
Authorizer:M. Carrano, R. Butler, P. Mannion, J. Alroy, R. Benson Enterer:M. Carrano, R. Benson, P. Mannion, R. Butler, E. Leckey
Modifier:M. Carrano Research group:vertebrate
Created:2002-04-25 13:34:17 Last modified:2025-02-22 15:12:02
Access level:the public Released:2002-04-25 13:34:17
Creative Commons license:CC0
Reference information

Primary reference:
9899. Z. Kielan-Jaworowska and R. Barsbold. 1972. Narrative of the Polish-Mongolian Palaeontological Expeditions 1967-1971. Palaeontologia Polonica 27:5-136 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

38117 V. R. Alifanov. 1996. Lizards of the families Priscagamidae and Hoplocercidae (Sauria, Iguania): phylogenetic position and new representatives from the Late Cretaceous of Mongolia. Paleontological Journal 30(4):466-483 [R. Benson/R. Benson]
38112 V. R. Alifanov. 2000. The fossil record of Cretaceous lizards from Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 368-389 [R. Benson/R. Benson/M. Carrano]
11744ETE V. R. Alifanov. 2003. Two new dinosaurs of the infraorder Neoceratopsia (Ornithischia) from the Upper Cretaceous of the Nemegt Depression, Mongolian People's Republic. Paleontological Journal 37(5):524-534 [M. Carrano/M. Carrano/M. Carrano]
14413ETE M. Borsuk-Bialynicka. 1985. Carolinidae, a new family of xenosaurid-like lizards from the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica 30(3-4):151-176 [M. Carrano/M. Carrano/M. Carrano]
37983 M. Borsuk-Bialynicka. 1988. Globaura venusta gen. et sp. n. and Eoxanta lacertifrons gen. et sp. n. - non-teiid lacertoids from the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica 33(3):211-248 [R. Benson/R. Benson]
14417ETE M. Borsuk-Bialynicka and S. M. Moody. 1984. Priscagaminae, a new subfamily of the Agamidae (Sauria) from the Late Cretaceous of the Gobi Desert. Acta Palaeontologica Polonica 29:51-81 [M. Carrano/M. Carrano/M. Carrano]
14221ETE K. Carpenter and K. Alf. 1994. Global distribution of dinosaur eggs, nests, and babies. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 13-30 [M. Carrano/M. Carrano/M. Carrano]
55988 A. Elzanowski. 1981. Emrbyonic bird skeletons from the Late Cretaceous of Mongolia. Palaeontologia Polonica 42:147-179 [M. Carrano/M. Carrano]
197345% 54920K. Gao and M. A. Norell. 2000. Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert. Bulletin of the American Museum of Natural History 249:1-118 [J. Alroy/E. Leckey/R. Benson]
10453 R. Gradzinski and T. Jerzykiewicz. 1972. Additional geographical and geological data from the Polish-Mongolian Palaeontological Expeditions. Palaeontologia Polonica 27:17-306 [M. Carrano/M. Carrano/M. Carrano]
62116 R. Gradzinski and T. Jerzykiewicz. 1974. Sedimentation of the Barun Goyot Formation. Palaeontologica Polonica 30:111-146 [M. Uhen/F. Villegas-Garin]
59136 R. Gradzinski, Z. Kielan-Jaworowska, and T. Maryanska. 1977. Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions. Acta Geologica Polonica 27(3):281-318 [M. Carrano/M. Carrano]
42976 J. H. Hurum and Z. Kielan−Jaworowska. 2008. Postcranial skeleton of a Cretaceous multituberculate mammal Catopsbaatar. Acta Palaeontologica Polonica 53(4):545-566 [P. Mannion/P. Mannion]
77615% 37520Z. Kielan-Jaworowska. 1974. Multituberculate succession in the Late Cretaceous of the Gobi Desert (Mongolia). Palaeontologia Polonica 30:23-44 [J. Alroy/E. Leckey/E. Leckey]
63572 Z. Kielan-Jaworowska. 1984. A caccia di dinosauri nel deserto di Gobi [Hunting dinosaurs in the Gobi Desert]. In G. M. Ronzoni, M. L. Greggio, & T. B. Guarinoni (eds.), Sulle Orme dei Dinosauri 87-109 [M. Carrano/M. Carrano]
43985 Z. Kielan-Jaworowska, J. H. Hurum, P. J. Currie and R. Barsbold. 2002. New data on anatomy of the Late Cretaceous multituberculate mammal Catopsbaatar. Acta Palaeontologica Polonica 47(3):557-560 [P. Mannion/P. Mannion]
38909 Z. Kielan-Jaworowska, J. H. Hurum, and D. Badamgarav. 2003. An extended range of the multituberculate Kryptobaatar and the distribution of mammals in the Upper Cretaceous of the Gobi Desert. Acta Palaeontologica Polonica 48(2):273-278 [R. Benson/R. Benson]
29707ETE M. Kundrát and J. Janácek. 2007. Cranial pneumatization and auditory perceptions of the oviraptorid dinosaur Conchoraptor gracilis (Theropoda, Maniraptora) from the Late Cretaceous of Mongolia. Naturwissenschaften 94:769-778 [M. Carrano/M. Carrano/M. Carrano]
18352ETE E. N. Kurochkin. 2000. Mesozoic birds of Mongolia and the former USSR. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 533-559 [M. Carrano/M. Carrano/M. Carrano]
60485% 6700T. Maryanska and H. Osmólska. 1975. Protoceratopsidae (Dinosauria) of Asia. Palaeontologica Polonica 33:133-181 [M. Carrano/M. Carrano/M. Carrano]
14217ETE K. Mikhailov, K. Sabath, and S. Kurzanov. 1994. Eggs and nests from the Cretaceous of Mongolia. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 88-115 [M. Carrano/M. Carrano/M. Carrano]
41004 K. Sabath. 1991. Upper Cretaceous amniotic eggs from the Gobi Desert. Acta Palaeontologica Polonica 36(2):151-192 [M. Carrano/M. Carrano]
10460 A. Sulimski. 1975. Macrocephalosauridae and Polyglyphanodontidae (Sauria) from the Late Cretaceous of Mongolia. Palaeontologia Polonica 33:25-102 [M. Carrano/M. Carrano/M. Carrano]