Woodring Loc. 176a - Margarita [Gatun Fm]: Tortonian, Panama
collected by Olsson 1918

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda - Heterostropha - Architectonicidae
Architectonica (Architectonica) nobilis (Röding 1798)
Woodring 1959 R category
original and current combination Architectonica nobilis
SUBSPECIES: Architectonica (Architectonica) nobilis nobilis
Gastropoda - Naticidae
Natica (Naticarius) stenopa
original and current combination Naticarius
Natica (Natica) bolus Brown and Pilsbry 1913
Stigmaulax guppiana (Toula 1909)
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria (Cancellaria) epistomifera (Guppy 1876)
Woodring 1970 R category
recombined as Bivetiella epistomifera
SUBSPECIES: Cancellaria (Cancellaria) epistomifera dariena
Gastropoda - Neogastropoda - Conidae
Woodring 1970 R category
Conus multiliratus (Olsson 1922)
Woodring 1970 R category
recombined as Conasprelloides multiliratus
SUBSPECIES: Conus multiliratus multiliratus
Gastropoda - Neogastropoda - Turridae
Pleurolira (Polystira) tenagos Gardner 1947
Woodring 1970 R category
    = Polystira sp. Woodring 1928
Todd and Johnson 2013
Gastropoda - Neogastropoda - Drilliidae
Crassispira (Hindsiclava) consors (Sowerby 1850)
Woodring 1970 R category
recombined as Hindsiclava consors
SUBSPECIES: Crassispira (Hindsiclava) consors consors
Gastropoda - Neogastropoda - Terebridae
Terebra (Oreoterebra) subsulcifera (Brown and Pilsbry 1911)
Woodring 1970 R category
Strioterebrum spiriferum (Dall 1895)
Woodring 1970 R category
Gastropoda - Neogastropoda - Turbinellidae
Xancus validus (Sowerby 1850)
Woodring 1964 R category
recombined as Turbinella valida
SUBSPECIES: Xancus validus falconensis
Gastropoda - Neogastropoda - Muricidae
Typhis (Pilsbrytyphis) woodringi Gertman 1969
Woodring 1973 R category
Typhis (Pilsbrytyphis) gabbi in Woodring (1959); Typhis (Pilsbrytyphis) woodringi in Woodring (1973)
Typhis (Pilsbrytyphis) gabbi (Brown and Pilsbry 1911)
Woodring 1959 R category
recombined as Siphonochelus (Pilsbrytyphis) gabbi
Gastropoda - Neogastropoda - Olividae
Oliva (Oliva) gatunensis (Toula 1908)
Woodring 1964 C category
original and current combination Oliva gatunensis
Gastropoda - Neogastropoda - Columbellidae
Strombina (Strombina) lessepsiana (Brown and Pilsbry 1911)
Woodring 1964 F category
Gastropoda - Neogastropoda - Buccinidae
Antillophos (Ripophos) metuloides (Dall 1896)
Woodring 1964 R category
Metula gabbi Brown and Pilsbry 1911
Woodring 1964 R category
Gastropoda - Personidae
Distorsio (Rhysema) decussata (Valenciennes 1832)
Woodring 1959 R category
recombined as Distorsio decussata
SUBSPECIES: Distorsio (Rhysema) decussata gatunensis
Scaphopoda - Dentaliida - Dentaliidae
Dentalium (Dentalium) armillatum
Woodring 1973 R category
SUBSPECIES: Dentalium (Dentalium) armillatum armillatum
Bivalvia - Cardiida - Chamidae
Arcinella arcinella (Linnaeus 1767)
Woodring 1982 R category
Bivalvia - Cardiida - Veneridae
Agiopoma (Pitarella) gatunensis (Dall 1903)
Woodring 1982 C category
recombined as Callocardia (Agriopoma) gatunensis
Bivalvia - Cardiida - Tellinidae
Tellina sp. Linnaeus 1758
Woodring 1982 R category
Psammacoma gatunensis (Toula 1909)
Woodring 1982 R category
Bivalvia - Lucinida - Lucinidae
Lucina (Lepilucina) gratis Olsson 1964
Woodring 1982 R category
Bivalvia - Pectinida - Pectinidae
Lepidopecten scissuratus (Dall 1898)
Woodring 1982 F category
Bivalvia - Ostreida - Ostreidae
Ostrea equestris Say 1834
Woodring 1982 R category
SUBSPECIES: Ostrea equestris heothina
Bivalvia - Arcida - Glycymerididae
Glycymeris (Tucetona) secticostata
Woodring 1973 C category
original and current combination Tucetona
SUBSPECIES: Glycymeris (Tucetona) secticostata schencki
Bivalvia - Arcida - Arcidae
Anadara (Tosarca) veatchi (Olsson 1922)
Woodring 1973 F category
SUBSPECIES: Anadara (Tosarca) veatchi veatchi
Anadara (Rasia) actinophora (Dall 1898)
Woodring 1973 R category
SUBSPECIES: Anadara (Rasia) actinophora actinophora
Anadara (Rasia) dariensis (Brown and Pilsbry 1911)
Woodring 1973 R category
SUBSPECIES: Anadara (Rasia) dariensis dariensis
see common names

Geography
Country:Panama State/province:Colon
Coordinates: 9.3° North, 79.9° West (view map)
Paleocoordinates:9.1° North, 78.5° West
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Tortonian 10 m.y. bin:Cenozoic 6
Key time interval:Tortonian
Age range of interval:11.63000 - 7.24600 m.y. ago
Age estimate:9.5 Ma (other)
Stratigraphy
Formation:Gatun Member:upper Gatun
Stratigraphic resolution:bed
Stratigraphy comments: LITHOSTRATIGRAPHY: From the upper part of the Gatun Formation. The Gatun Formation is subdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Late Miocene, on the basis of microfossil biostratigraphy (see Collins and Coates, 1999), refined by enterer to Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 9.5 Ma (Hendy, unpublished data).
Lithology and environment
Primary lithology: poorly lithified siltstone
Secondary lithology: poorly lithified silty sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: LITHOLOGY: Siltstone and silty sandstone.
Environment:marine indet.
Geology comments: No specific environmental data presented.
Taphonomy
Modes of preservation:body,original aragonite
Size of fossils:macrofossils
Collection methods and comments
Collection methods:surface (float),surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Museum repositories:USNM
Collectors:Olsson Collection dates:1918
Collection method comments: COLLECTOR: Olsson. REPOSITORY: USNM.
Taxonomic list comments:COVERAGE: Exhaustive for gastropods and bivalves. NOMENCLATURE: Authoritative publications, with modern nomenclature, and species-resolution identifications.
Metadata
Also known as:USGS 8409; WG176a
Database number:145342
Authorizer:A. Hendy Enterer:A. Hendy
Research group:marine invertebrate
Created:2013-06-01 00:29:35 Last modified:2013-11-25 13:37:23
Access level:the public Released:2013-06-01 00:29:35
Creative Commons license:CC BY
Reference information

Primary reference:

11306. W. P. Woodring. 1957. Geology and Paleontology of Canal Zone and adjoining parts of Panama. United States Geological Survey Professional Paper 306(A) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

48647 J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy]
11372 W. P. Woodring. 1959. Geology and paleontology of Canal Zone and adjoining parts of Panama: Description of Tertiary mollusks (gastropods: Vermetidae to Thaididae). United States Geological Survey Professional Paper 306(B):147-239 [A. Miller/A. Hendy/A. Hendy]
13232 W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy]
13236 W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy]
13235 W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy]
13359 W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]