Cerro Zeballos : Tortonian, Argentina
collected by Brandoni et al. 2019

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Notoungulata - Interatheriidae
Protypotherium cf. endiadys (Roth 1898)
Vera et al. 2019
LIEB-PV 7082
Mammalia - Notoungulata
Hemihegetotherium tantillum n. sp. Vera 2019
Vera 2019
LIEB-PV 7181 (type)
Mammalia - Nothrotheriidae
"Xyophorus" sp. Ameghino 1887
LIEB-PV 5129
    = Mcdonaldocnus bondesioi Scillato-Yané 1979
Gaudin et al. 2022
LIEB PV 5129, right mandibular fragment
Mammalia - Megatheriidae
Megathericulus patagonicus Ameghino 1904
Brandoni et al. 2019
LIEB-PV 6153 - a right astragalus
see common names

Geography
Country:Argentina State/province:Chubut
Coordinates: 42.6° South, 70.3° West (view map)
Paleocoordinates:42.9° South, 67.7° West
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period: Neogene Epoch: Miocene
Stage: Tortonian 10 m.y. bin: Cenozoic 6
Key time interval: Tortonian
Age range of interval: 11.63 - 7.246 m.y. ago
Stratigraphy
Formation:Collón Curá
Stratigraphy comments: Brandoni et al. 2017: The new specimen presented here as ‘Xyophorus’ sp. (LIEB PV 5129) has a DL/TRL index of ca. 14, this value being larger than the index for ‘X.’ villarroeli (12.07) from the Mauri Formation (ca. 10.3 Ma), but smaller than the value for ‘X.’ bondesioi (16.45) from Arroyo Chasicó Formation (ca. 10–8.7 Ma) (Table 1, Figure 6). Following the relationship between DL/TRL index and age proposed by Brandoni (2014), where the value of the index would decrease with the age of the bearing sediments, the DL/TRL index of ca. 14 for ‘Xyophorus’ sp. (LIEB PV 5129) would suggest a Tortonian age (11.63–7.24 Ma) for the deposits of Collón Curá Formation at Cerro Zeballos (Figure 6), which results in a ‘younger age’ compared to the middle Miocene age traditionally accepted for the Collón Curá Formation with the Colloncuran fauna sensu stricto. Although no absolute ages for Cerro Zeballos are available yet, the geographic proximity of Cerro Zeballos to Cushamen River (with levels dated in ca. 11.2 Ma) supports the possibility that the sediments yielding specimen LIEB PV 5129 are younger than those from those associated to the ignimbrite that carry the Colloncuran fauna sensu stricto, so the tenta- tive Tortonian age (late Miocene) indicated by the presence of ‘Xyophorus’ sp (LIEB PV 5129).
Brandoni et al. 2019: there are no absolute dates for Cerro Zeballos, the chronological location of this locality in Fig. 6a is relative and based on the study of Brandoni et al. (2017) that indicated an early Tortonian age (its position could change if absolute dates appear).
Vera, 2019: he Collón Curá Formation (Roth, 1899; Yrigoyen, 1969) is widely distributed in western Patagonia, from Neuquén Province (e.g., Estancia Collón Curá, Cañadón del Tordillo) in the North, to Río Negro (e.g., Comallo, Estancia El Criado) and Chubut (e.g., Río Chico, Cerro Zeballos) provinces in the South, ranging in age from approximately 16 Ma to 11 Ma (Burdigalian–Tortonian; Marshall et al., 1977; Mazzoni and Benvenuto, 1990; Bilmes et al., 2013; Ramos et al., 2015; Brandoni et al., 2017).

Vera, et al. 2018: Several absolute ages for the Coll?on Cur?a Formation indicate that this unit ranges from approximately 16Ma Miocene; Marshall et al. 1977; Mazzoni & Benvenuto to 11Ma (Burdigalian?Tortonian; early to late 1990; Bilmes et al. 2013;Ramos et al. 2015). Unfortunately, there is no absolute age for the Coll?on Cur?a Formation at Cerro Zeballos, but some of the recovered mammal taxa differ from the typical Colloncuran (Martin & Tejedor 2007; Gonz?alez Ruiz et al. 2011; Martin et al. 2016)and suggest a Tortonian age (late Miocene; Brandoni et al. 2017 and references cited therein)
Lithology and environment
Primary lithology:tabular,concretionary,tuffaceous,yellow lithified sandstone
Secondary lithology: lithified "limestone"
Includes fossils?Y
Lithology description: Brandoni et al. 2019: The analyzed section starts with a 15 m thick of massive ocher or yellowish white tuffaceous sandstone, in general very friable and powdery. Tabular bodies with concentrations of reddish concretions composed by tuffitic material frequently appear. These levels show the presence of roots and scarce paleosol development. Fossil vertebrates are abundant throughout the section, even inside the concretions. Specimen LIEB-PV 6153 was collected in the middle of this 15 m section (Fig. 2b). The profile continues with 3 m thick of greenish tuffaceous limestones with intercalated levels of ocher tuffaceous and massive fine sandstones. Above this, the profile continues with volcaniclastic to epiclastic deposits of claystone, mudstone, and grainstones, with some pyroclastic subordinated levels intercalated (Brandoni et al. 2017). Brandoni et al. 2017: The described profile (Figure 2) starts with a section of ca. 15 m of massive ocher or yellowish white tuffaceous sandstone, in general very friable and powdery, forming the typical badlands landscape. In all the profile, levels with concentrations of reddish concretions composed by tuffitic material frequently appears (Tf in Figure 2), and because they are more resistant to weathering they form steps in the profile. These levels with concretions and the presence of roots could indicate pedogenic processes. In all the section fossil vertebrates are abundant, even inside these con- cretions. Specimen LIEB PV 5129 was collected in the middle of this 15 m section. The profile continues, in paracomformity, with ca. 3 m of greenish tuffaceous mudstone with intercalated levels of ocher tuffaceous fine sandstones with massive structures (T in Figure 2). Above this, the profile continues with epiclastic deposits of claystone, mudstone, and grainstones, with some pyroclastic subordinated levels intercalated.
Environment:alluvial fan
Geology comments: Brandoni et al. 2019: The presence of mainly massive and tabular sandstones with scarce paleosol development indicates a shallow and low energy system, probably related with alluvial to fluvial plains located close to shallow lagoons or lakes (Brandoni et al. 2017).
Brandoni et al. 2017: According to Bilmes et al. (2014) the Collón Curá Formation, near this area, is the result of the conformation of isolated depocenters in which ephemeral and deep lacustrine systems were developed. The described section has no structures that could indicate the deposition agent, although the tuffaceous material and the presence of roots suggests a shallow and low energy system of lagoons or lakes, and a floodplain with shallow and low energy fluvial courses, probably with periods of aerial exposure.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collectors:Brandoni et al. Collection dates:2019
Collection method comments: Brandoni et al. 2019: LEIB-PV 6153 (a complete right astragalus) is housed in Laboratorio de Investigaciones en Evolución y Biodiversidad (PV, Paleovertebrados), Esquel, Argentina.
Brandoni et al. 2017: LIEB PV 5129 is housed in Laboratorio de Investigaciones en Evolución y Biodiversidad (-PV, Paleovertebrados), Esquel, Argentina.
Vera, 2019: The fossil specimens described below were recovered by the CIEMEP paleontology team during several field seasons (2003–2018) to Cerro Zeballos (Chubut Province; Fig. 1) and they are sheltered in the vertebrate paleontology collection of the Laboratorio de Investigaciones en Evolución y Biodiversidad (LIEB), Esquel, Chubut, Argentina.
Metadata
Database number:201907
Authorizer:P. Mannion Enterer:G. Varnham, M. Kouvari
Modifier:M. Kouvari Research group:vertebrate
Created:2019-06-09 07:46:33 Last modified:2020-09-20 08:13:59
Access level:the public Released:2019-06-09 07:46:33
Creative Commons license:CC BY
Reference information

Primary reference:

69184. D. Brandoni, L. González Ruiz, A. Reato and G. Martin. 2017. Chronological implications of the nothrotheriid ‘Xyophorus’ (Mammalia, Xenarthra) from the Collón Curá Formation (Miocene of Patagonia, Argentina). Historical Biology [P. Mannion/G. Varnham/G. Varnham]

Secondary references:

69135 D. Brandoni, L. González Ruiz, and J. Bucher. 2019. Evolutive Implications of Megathericulus patagonicus (Xenarthra, Megatheriinae) from the Miocene of Patagonia Argentina. Journal of Mammalian Evolution 1-16 [P. Mannion/G. Varnham]
81883 T. J. Gaudin, A. Boscaini, B. M. Quispe, R. A. Flores, M. Fernández-Monescillo, L. Marivaux, P. Antoine, P. Münch, and F. Pujos. 2022. Recognition of a new nothrotheriid genus (Mammalia, Folivora) from the early late Miocene of Achiri (Bolivia) and the taxonomic status of the genus Xyophorus. Historical Biology [P. Mannion/G. Varnham]
69350 B. Vera. 2019. A new species and the record of Hemihegetotherium (Notoungulata, T Hegetotheriidae) in the Middle to Late Miocene of Patagonia, Argentina. Journal of South American Earth Sciences 93:23-35 [P. Mannion/G. Varnham]
70219 B. Vera, L. Gonzalez-Ruiz, N. Novo, G. Martin, A. Reato and M. F. Tejedor. 2019. The Interatheriinae (Mammalia, Notoungulata) of the Friasian sensu stricto and Mayoan (middle to late Miocene), and the fossils from Cerro Zeballos, Patagonia, Argentina. Journal of Systematic Palaeontology 17(13):1143-1163 [P. Mannion/M. Kouvari/K. Pino]