Leeukloof: Wuchiapingian, South Africa

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Scylacognathus major n. sp. Broom 1935
Broom 1935 1 individual
TM P256
Cynarioides grimbeeki n. sp. Broom 1935
Sigogneau-Russell 1989 1 individual
TMP 246, crushed skull and mandible
Gorgonopidae
Aelurosaurus sp. Owen 1881
Sigogneau-Russell 1989 1 individual
SAM-PK-10678
Galerhinus polyodon n. gen., n. sp. Broom 1935
1 individual
TM P341, snout with part of lower jaw
Aelurosaurus brevirostris n. sp. Broom 1948
Broom 1948 1 individual
Specimen now lost
Anomodontia - Cryptodontidae
Tropidostoma microtrema (Seeley 1889)
Dicynodon acutirostris n. sp. Broom 1935
Broom 1935 1 individual
TM 250 , partial skull
Dicynodon validus n. sp. Broom 1935
Broom 1935 1 individual
TM 252, a distorted snout and anterior portion of the lower jaws
Anomodontia
Pristerodon indet. Huxley 1868
Emydopsis microdon n. sp. Broom 1936
TM 269, a skull
Eumantellia mirus n. gen., n. sp. Broom 1935
1 individual
TM 248, a rather badly weathered anterior part of skull
Palemydops minor n. sp. Broom 1935
Lefebvre and Gutierrez-Marco 2003
TM 247, an imperfect skull
Anomodontia - Pristerodontidae
Emyduranus gracilis n. sp. Broom 1935
Broom 1935
TM 243, a skull
Anomodontia
Diictodon sp. Broom 1913
Smith 1993 12 individuals
Dicynodon grimbeeki n. sp. Broom 1935
Broom 1935
TM 253, a complete skull
Reptilia
Pareiasaurus serridens Owen
see common names

Geography
Country:South Africa State/province:Western Cape County:Beaufort West
Coordinates: 32.1° South, 22.0° East (view map)
Paleocoordinates:50.4° South, 21.0° West (Wright 2013)
Geographic resolution:outcrop
Time
Period: Permian Epoch: Lopingian
Stage: Wuchiapingian 10 m.y. bin: Permian 4
*International age/stage:Kungurian - Olenekian
Key time interval: Wuchiapingian Zone:  Tropidostoma Assemblage Zone
Age range of interval: 259.51 - 254.14 m.y. ago
Age estimate: 259.26 to 256.25 Ma (U/Pb)
* legacy (obsolete) database fields
Stratigraphy
Geological group:Beaufort Formation:Teekloof
Stratigraphy comments: Originally entered as Kungurian to Olenekian

Direct date for the Tropidostoma Assemblage Zone from Rubidge et al., 2013

Lithology and environment
Primary lithology: sandstone
Secondary lithology: claystone
Lithology description: ETE rock type adj: 0, ETE lithology comments: .
Environment:crevasse splay
Geology comments: ETE sed env 1: fluvial, ETE sed env 2: crev_splay ETE event: flood, ETE env comment: 0
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collection method comments: ETE Size: .; ETE reference list: 478, ; ETE museum list:
Metadata
Also known as:Leeupoort, Leeuwkloof, ETE Locality 1536
Database number:22661
Authorizer:A. Behrensmeyer, R. Butler Enterer:A. Behrensmeyer, R. Butler, J. Benito Moreno
Modifier:J. Benito Moreno Research group:vertebrate
Created:2001-04-17 01:00:00 Last modified:2025-02-22 15:12:02
Access level:the public Released:2001-04-17 01:00:00
Creative Commons license:CC0
Reference information

Primary reference:

27216. J. W. Kitching. 1977. The distribution of the Karroo vertebrate fauna. Memoirs of the Bernard Price Institute for Palaeontological Research 1:1-131 [H. Sims/J. Camp]

Secondary references:

56911 R. Broom. 1935. On some new genera and species of Karroo fossil reptiles. Annals of the Transvaal Museum 18:55-72 [R. Butler/J. Benito Moreno]
57014 R. Broom. 1948. A Contribution to our Knowledge of the Vertebrates of the Karroo Beds of South Africa. Transactions of the Royal Society of Edinburgh 61(2):577-629 [R. Butler/J. Benito Moreno/J. Benito Moreno]
51786 C. F. Kammerer, K. D. Angielczyk, and J. Frobisch. 2011. A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia) and its implications for dicynodont phylogeny, biogeography, and biostratigraphy. Journal of Vertebrate Paleontology 31(Sup. to 6):1-158 [M. Uhen/M. Uhen/J. Benito Moreno]
65911 B. Lefebvre and J. C. Gutierrez-Marco. 2003. New Ordovician mitrocystitidan mitrates (Echinodermata, Stylophora) from the Central Iberian zone (Spain). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 227(1):39-61 [P. Novack-Gottshall/P. Novack-Gottshall]
57886 D. S. Sigogneau-Russell. 1989. Theriodontia I: Phthinosuchia, Biarmosuchia, Eotitanosuchia, Gorgonopsia. Encyclopedia of Paleoherpetology [R. Butler/R. Butler/R. Butler]
5708ETE 1464R. M. H. Smith. 1993. Vertebrate taphonomy of late Permian floodplain deposits in the Southwestern Karoo Basin of South Africa. Palaios 8:45-67 [A. Behrensmeyer/A. Behrensmeyer/M. Kosnik]