PBDB Collection

Vallipón, Castellote: Early/Lower Barremian, Spain
collected 1995

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Amphibia - Temnospondyli

	Anura indet. (Fischer von Waldheim 1813)	Ruiz-Omeñaca and Canudo 2001
Reptilia - Dinosauria

	Dinosauria indet. Owen 1842
Reptilia

	? Stegosauria indet. Marsh 1877	Ruiz-Omeñaca and Canudo 2001

	Ornithopoda indet. Marsh 1881	Ruiz-Omeñaca and Canudo 2001
Reptilia - Hypsilophodontidae

	Hypsilophodontidae indet. Dollo 1882		2 specimens
	MPZ 96/57
Reptilia

	cf. Hypsilophodon sp. Huxley 1870	Ruiz-Omeñaca and Canudo 2001

	Iguanodon sp. Mantell 1825		3 specimens
	MPZ 96/50–52
	= Iguanodontoidea indet. Hay 1902	Gasca et al. 2008
	morphotype B

	Iguanodontidae indet. Bonaparte 1850	Ruiz-Omeñaca and Canudo 2001
	= Iguanodontoidea indet. Hay 1902	Gasca et al. 2008
	morphotype B
Reptilia - Theropoda

	Theropoda indet. Marsh 1881	Ruiz-Omeñaca and Canudo 2001
Reptilia

	Coelurosauria indet. Huene 1914	Ruiz-Omeñaca and Canudo 2001
Reptilia - Dromaeosauridae

	Dromaeosauridae indet. Colbert and Russell 1969	Canudo et al. 1997	1 specimen
	MPZ 96/100
Reptilia - Spinosauridae

	Baryonychidae indet. Charig and Milner 1986	Ruiz-Omeñaca and Canudo 2001
	spelled with current rank as Baryonychinae
Reptilia

	Sauropoda indet. Marsh 1878	Ruiz-Omeñaca and Canudo 2001
	= Pleurocoelus valdensis Lydekker 1889	Ruiz-Omeñaca and Canudo 2005
	nomen dubium belonging to Sauropoda

	Titanosauriformes indet. Salgado et al. 1997	Barco et al. 1999	1 specimen
	MPZ 99/672

	Pterodactyloidea informal indet. A Plieninger 1901	Ruiz-Omeñaca et al. 1998	21 specimens
	MPZ 96/59, 98/39 to 98/48; 21 teeth

	Pterodactyloidea informal indet. B Plieninger 1901	Ruiz-Omeñaca et al. 1998	1 specimen
	MPZ 96/58
Reptilia - Goniopholididae

	Goniopholis sp. Owen 1842	Ruiz-Omeñaca and Canudo 2001
Reptilia - Crocodylia

	Crocodylia indet. (Owen 1842)
Reptilia

	Theriosuchus sp. Owen 1878	Ruiz-Omeñaca and Canudo 2001
	= aff. Theriosuchus sp. Owen 1878	Tennant et al. 2016

	Bernissartia sp. Dollo 1883	Ruiz-Omeñaca and Canudo 2001
Reptilia - Plesiosauria

	Plesiosauria indet. de Blainville 1835	Ruiz-Omeñaca and Canudo 2001
Reptilia - Paramacellodidae

	Paramacellodidae indet. Estes 1983	Ruiz-Omeñaca and Canudo 2001
Reptilia - Testudines

	Cryptodira indet. Cope 1868	Ruiz-Omeñaca and Canudo 2001
Reptilia - Testudines - Dortokidae

	Dortokidae indet. Lapparent de Broin and Murelaga 1996	Ruiz-Omeñaca and Canudo 2001
	= ? Dortoka sp. Lapparent de Broin and Murelaga 1996	Lapparent de Broin et al. 2004
	V 9326 (right posterior pleural)
	= Dortokidae indet. Lapparent de Broin and Murelaga 1996	Cadena and Joyce 2015
	p.17
Docodonta

	Docodonta indet. Kretzoi 1946	Cuenca-Bescós and Canudo 2003
Mammalia - Multituberculata - Paulchoffatiidae

	Paulchoffatiinae indet. Hahn 1971	Cuenca-Bescós et al. 1996	2 specimens
	MPZ 96/174, 96/176
Mammalia - Multituberculata - Pinheirodontidae

	Eobaatarinae indet. (Kielan-Jaworowska et al. 1987)	Cuenca-Bescós et al. 1996	2 specimens
	MPZ 96/165, 96/175
	= ? Lavocatia sp. Canudo and Cuenca-Bescós 1996	Ruiz-Omeñaca and Canudo 2001
	'Some isolated teeth (I2, anterior upper premolars, and upper molars) figured and briefly described by Cuenca-Bescós et al. (1995) from the Barremian of Spain and assigned to Eobaatarinae indet. and Paulchoffatinae indet. might also belong to Lavocatia'
Mammalia - Multituberculata - Eobaataridae

	Eobaatarinae indet. (Kielan-Jaworowska et al. 1987)	Cuenca-Bescós et al. 1996	8 specimens
	original and current combination Eobaataridae
	MPZ 96/166–173

	Eobaatarinae indet. (Kielan-Jaworowska et al. 1987)	Cuenca-Bescós et al. 1996	9 specimens
	MPZ 96/156–159, MPZ2010/859–862; Three P1/3s... a ?P5 of Eobaatar may also belong to this species... other teeth such as the three upper incisors and upper and lower first and second molars may also belong to this species of Eobaatar
	= Eobaatar sp. Kielan-Jaworowska et al. 1987	Badiola et al. 2012

	Eobaatarinae indet. (Kielan-Jaworowska et al. 1987)	Cuenca-Bescós et al. 1996	7 specimens
	MPZ 96/160–164
	= Plagiaulacoidea indet. Ameghino 1889	Ruiz-Omeñaca and Canudo 2001
	= ? Eobaatar pajaronensis Hahn and Hahn 2001	Badiola et al. 2012
	MPZ 2010/863, 2010/865; Seven P1/3s resemble those of Eobaatar? pajronensis
Mammalia - Symmetrodonta

	Symmetrodonta indet. Simpson 1925	Cuenca-Bescós et al. 1996	1 specimen
	MPZ 96/180
Mammalia

	? Theria indet. Parker and Haswell 1897	Cuenca-Bescós et al. 1996	3 specimens
	MPZ 96/177–179
Mammalia - Eutriconodonta - Gobiconodontidae

	Gobiconodontidae indet. Jenkins and Schaff 1988	Cuenca-Bescós and Canudo 2003
	= Gobiconodon sp. Trofimov 1978	Ruiz-Omeñaca and Canudo 2001
	"sp. nov"
Actinopteri

	Osteichthyes indet. Huxley 1880
	= Holostei indet. Müller 1846	Ruiz-Omeñaca and Canudo 2001
Actinopteri - Amiiformes - Amiidae

	Amiidae indet. Huxley 1861	Ruiz-Omeñaca and Canudo 2001
Actinopteri - Lepisosteiformes

	Lepidotes sp. Agassiz 1832	Ruiz-Omeñaca and Canudo 2001
Actinopteri - Pycnodontiformes - Pycnodontidae

	Pycnodontidae indet. (Agassiz 1833)	Ruiz-Omeñaca and Canudo 2001
Chondrichthyes

	Chondrichthyes indet. Huxley 1880
Chondrichthyes - Synechodontiformes - Orthacodontidae

	Orthacodontidae indet. de Beaumont 1960	Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Rajiformes

	Pseudohypolophus mcnultyi (Thurmond 1971)	Ruiz-Omeñaca and Canudo 2001
	misspelled "mcnultyui"
Chondrichthyes - Rajiformes - Rhinobatidae

	Rhinobatos cf. halteri Biddle and Landemaine 1988	Ruiz-Omeñaca and Canudo 2001
	recombined as Spathobatis halteri
Chondrichthyes - Lamniformes - Odontaspididae

	Odontaspididae indet. Müller and Henle 1839	Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Lamniformes - Eoptolamnidae

	Protolamna cf. sokolovi Cappetta 1980	Ruiz-Omeñaca and Canudo 2001	50 specimens
	= Eoptolamna eccentrolopha n. gen., n. sp. Kriwet et al. 2008	Kriwet et al. 2008
Chondrichthyes - Hybodontiformes - Lonchidiidae

	Lissodus aff. crenulatus (Patterson 1966)	Ruiz-Omeñaca and Canudo 2001
	recombined as Lonchidion crenulatum
Chondrichthyes - Hybodontiformes

	Lissodus sp. Brough 1935	Ruiz-Omeñaca and Canudo 2001

	Lissodus breve	Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Hybodontiformes - Polyacrodontidae

	Polyacrodus sp. Jaekel 1889	Ruiz-Omeñaca and Canudo 2001

	Polyacrodus brevicostatus (Patterson 1966)	Ruiz-Omeñaca and Canudo 2001

	? Hybodus parvidens Woodward 1916	Ruiz-Omeñaca and Canudo 2001
	recombined as Polyacrodus parvidens
Gastropoda

	Gastropoda indet. Cuvier 1795	Ruiz-Omeñaca et al. 2003
Bivalvia

	Bivalvia indet. Linnaeus 1758	Ruiz-Omeñaca et al. 2003
see common names

Geography

Country:	Spain	State/province:	Aragón	County:	Teruel
Coordinates:	40.8° North, 0.3° West (view map)
Paleocoordinates:	30.6° North, 9.7° East
Basis of coordinate:	estimated from map
Altitude:	559 meters
Geographic resolution:	local area

Time

Period:	Cretaceous	Epoch:	Early/Lower Cretaceous
Stage:	Barremian	10 m.y. bin:	Cretaceous 2
*Period:	Early/Lower Cretaceous	*Epoch:	Early/Lower Gallic
*International age/stage:	Late/Upper Barremian
Key time interval:	Early/Lower Barremian
Age range of interval:	125.77000 - 121.40000 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Formation:	Artoles
Stratigraphic resolution:	member
Stratigraphy comments: base of this formation (original name translates as "Artoles Marls and Limestones") Villanueva-Amadoz et al. 2015: The presence of cf. Arecipites microfoveolatus Ibrahim, Afropollis sp. and Stellatopollis barghoornii Doyle from the basal levels of Artoles Formation suggests a late Barremian age (Pons et al. 2013). The schizacean fern spore content supports a Barremian age. Bover-Arnal et al. 2016: Sr values from samples suggest a middle/late Early Barremian age for the lower-middle part of the Artoles Formation, and a Late Barremian age for the upper part. Previously the Barremian–Aptian boundary has been placed within the Artoles Formation, but late Barremian ammonites have been found in the overlying Morella Formation, ruling out an early Aptian age for the Artoles Formation.

Lithology and environment

Primary lithology:	lithified calcareous sandstone
Secondary lithology:	conglomerate
Includes fossils?	Y
Includes fossils?	Y
Lithology description: Quartitic sandstones and conglomerates. "The rock is dissolved with 10% formic acid" before screenwashing.
Environment:	estuary/bay
Geology comments: "deposited in an intertidal or subtidal environment very close [related[ to the coastline"

Taphonomy

Modes of preservation:	body,trace,coprolite
Degree of concentration:	concentrated
Size of fossils:	macrofossils,mesofossils
Preservation of anatomical detail:	variable
Articulated whole bodies:	none
Associated major elements:	none
Disassociated major elements:	many
Disassociated minor elements:	many
Fragmentation:	frequent
Bioerosion:	frequent
Feeding/predation traces:	gastric dissolution
Temporal resolution:	time-averaged

Collection methods and comments

Collection excludes:	some genera,species names
Collection methods:	bulk,chemical,sieve,field collection
Minimum sieve size:	0.500
Reason for describing collection:	taxonomic analysis
Collection dates:	1995
Collection method comments: Materials include bones, coprolites, and gastroliths. Some preparation using formic acid.

Metadata

Database number:	28196
Authorizer:	M. Carrano, E. Vlachos, R. Benson, P. Mannion	Enterer:	M. Carrano, E. Vlachos, K. Maguire, P. Mannion, R. Benson
Modifier:	G. Varnham	Research group:	vertebrate
Created:	2003-01-28 15:00:53	Last modified:	2022-03-01 04:15:18
Access level:	the public	Released:	2003-01-28 15:00:53
Creative Commons license:	CC BY

Reference information

Primary reference:

7706.

5% 3240

J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1996. Dientes de dinosaurios ornitópodos en el Barremiense superior de Castellote y Mas de las Matas (Teruel) [Ornithopod dinosaur teeth in the Upper Barremian of Castellote and Mas de las Matas (Teruel)]. Tomo Estraordinario, 125 Aniversario de la RSEHN. Real Sociedad Española de Historia Natural, Madrid 315-319 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

28224	ETE	A. Badiola, J. I. Canudo, and G. Cuenca-Bescós. 2007. Los fósiles de multituberculados del Hauteriviense y Barremiense de la Península Ibérica [The multituberculate fossils from the Hauterivian and Barremian of the Iberian Peninsula]. IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Salas de los Infantes, Spain 47-48 [M. Carrano/M. Carrano]
42710		A. Badiola, J. I. Canudo, and G. Cuenca-Bescós. 2012. New Early Cretaceous multituberculate mammals from the Iberian Peninsula. In P. Godefroit (ed.), Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems 409-433 [R. Benson/R. Benson]
17666	ETE	J. L. Barco, J. I. Canudo, J. I. Ruiz-Omeñaca and R. Royo-Torres. 1999. Bones, teeth and tracks: about sauropod dinosaur remains from Aragón (northeastern Spain). In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 22-23 [M. Carrano/M. Carrano/M. Carrano]
80368		T. Bover-Arnal, J. A. Moreno-Bedmar, G. Frijia, E. Pascual-Cebrian, and R. Salas. 2016. Chronostratigraphy of the Barremian-Early Albian of the Maestrat Basin (E Iberian Peninsula): integrating strontium-isotope stratigraphy and ammonoid biostratigraphy. Newsletters on Stratigraphy 49(1):41-68 [P. Mannion/G. Varnham/P. Mannion]
62214		E. Cadena and W. G. Joyce. 2015. A Review of the Fossil Record of Turtles of the Clades Platychelyidae and Dortokidae. Bulletin of the Peabody Museum of Natural History 56(1):3-20 [E. Vlachos/E. Vlachos/P. Mannion]
53564		J. I. Canudo, J. L. Barco, R. Royo-Torres and J. I. Ruiz-Omeñaca. 2001. Los saurópodos (Dinosauria) del Tithónico (Jurásico Superior) y del Cretácico de Aragón [The sauropoda (Dinosauria) from the Tithonian (Upper Jurassic) and Cretaceous of Aragón]. I Jornadas Internacionales sobre Paleontología de Dinosaurios y Su Entorno Actas:309-318 [M. Carrano/M. Carrano/M. Carrano]
14676	ETE	J. I. Canudo, G. Cuenca-Bescós, and J. I. Ruiz-Omeñaca. 1997. Dinosaurios dromaeosaúridos (Saurischia: Theropoda) en el Barremiense Superior (Cretácico Inferior) de Castellote, Teruel [Dromaeosaurid dinosaurs (Saurischia: Theropoda) in the upper Barremian (Lower Cretaceous) of Castellote, Teruel)]. Geogaceta 22:39-42 [M. Carrano/K. Maguire/M. Carrano]
17647	ETE	J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
41617		M. T. Carrano, R. B. J. Benson, and S. D. Sampson. 2012. The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology 10(2):211-300 [M. Carrano/M. Carrano]
54838		G. Cuenca-Bescós, J. I. Canudo, and J. I. Ruiz-Omeñaca. 1996. Los mamíferos del Barremiense superior (Cretácico inferior) de Vallipón (Castellote, Teruel) [The mammals of the upper Barremian (Lower Cretaceous) of Vallipón (Castellote, Teruel)]. Beca del Museo de Mas de las Matas Año 1995:105-137 [M. Carrano/M. Carrano]
14443	ETE	G. Cuenca-Bescós and J. I. Canudo. 2003. A new gobiconodontid mammal from the Early Cretaceous of Spain and its palaeogeographic implications. Acta Palaeontologica Polonica 48(4):575-582 [M. Carrano/M. Carrano/M. Carrano]
17652	ETE	G. Cuenca-Bescós and J. I. Canudo. 2004. Los mamíferos del Cretácico inferior de España [Lower Cretaceous mammals from Spain]. Geo-Temas 6(5):35-38 [M. Carrano/M. Carrano]
43435		V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2012. Juvenile and adult teeth of the titanosaurian dinosaur Lirainosaurus (Sauropoda) from the Late Cretaceous of Iberia. Geobios 45:265-274 [M. Carrano/M. Carrano]
31341	ETE	J. M. Gasca, J. I. Canudo, and M. Moreno-Azanza. 2008. Revisión de morfotipos dentales de los iguanodontoideos del Cretácico Inferior de Teruel [Revision of dental morphotypes of the iguanodontoids from the Lower Cretaceous of Teruel]. In J I Ruiz-Omeñaca, L Piñuela and J C García-Ramos (eds), XXIV Jornadas de la Sociedad Española de Paleontología, 15-18 October 2008, Museo del Jurásico de Asturias (MUJA), Colunga, Spain, Libro de Resúmenes 127-128 [M. Carrano/M. Carrano]
14365		Z. Kielan-Jaworowska, R. L. Cifelli, and Z.-X. Luo. 2004. Mammals from the age of dinosaurs: Origins, evolution, and structure. Columbia University Press, New York 1-630 [R. Whatley/R. Whatley/R. Whatley]
28298	ETE	J. Kriwet, S. Klug, J. I. Canudo and G. Cuenca-Bescós. 2008. A new Early Cretaceous lamniform shark (Chondrichthyes, Neoselachii). Zoological Journal of the Linnean Society 154:278-290 [M. Carrano/M. Carrano]
44196		F. d. Lapparent de Broin, X. Murelaga Bereikua, and V. Codrea. 2004. Presence of Dortokidae (Chelonii, Pleurodira) in the earliest Tertiary of the Jibou Formation, Romania: paleobiogeographical implications. Acta Palaeontologica Romaniae 4:203-215 [P. Mannion/P. Mannion]
17663	ETE	R. Royo-Torres, J. L. Barco, J. I. Canudo and J. I. Ruiz-Omeñaca. 2000. Descripción de un fragmento de costilla de Dinosauria (Saurópoda), del Barremiense superior del yacimiento de Vallipón (Castellote, Teruel) [Description of a dinosaur rib fragment (Sauropoda), from the upper Barremian of the Vallipón locality (Castellote, Teruel)]. Mas de las Matas 19:121-132 [M. Carrano/M. Carrano]
51920		J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1998. Primeros restos de reptiles voladores (Pterosauria: Pterodactyloidea) en el Barremiense superior (Cretacico inferior) de Vallipón (Castellote, Teruel) [First remains of flying reptiles (Pterosauria: Pterodactyloidea) in the upper Barremian (Lower Cretaceous) of Vallipón (Castellote, Teruel)]. Mas de las Matas 17:225-249 [M. Carrano/M. Carrano]
29947	ETE	J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1998. Sobre las especies de Iguanodon (Dinosauria, Ornithischia) encontradas en el Cretácico inferior de España [On the species of Iguanodon (Dinosauria, Ornithischia) found in the Lower Cretaceous of Spain]. Geogaceta 24:275-278 [M. Carrano/M. Carrano]
14802	ETE	J. I. Ruiz-Omeñaca and J. I. Canudo. 2001. Dos yacimientos excepcionales con vertebrados continentales del Barremiense (Cretácico Inferior) de Teruel: Vallipón y La Cantalera [Two exceptional localities with continental vertebrates in the Barremian (Lower Cretaceous) of Teruel: Vallipón and La Cantalera]. Naturaleza Aragonesa 7:8-18 [M. Carrano/M. Carrano]
55492		J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 2002. Los dinosaurios del Jurásico y del Cretácico de la Cordillera Ibérica aragonesa [The Jurassic and Cretaceous dinosaurs of the Aragonese Iberian Cordillera]. Actas del I Encuentro Nacional de Estudios sobre la Cordillera Ibérica (ciencias de la naturaleza) 119-130 [M. Carrano/M. Carrano]
17657	ETE	J. I. Ruiz-Omeñaca, J. M. Canudo, and G. Cuenca-Bescós. 2003. Vallipón, el yacimiento con vertebrados continentales del Cretácico Inferior más completo de Europe [Vallipón, the most complete locality with Lower Cretaceous continental vertebrates in Europe]. El Masino [M. Carrano/M. Carrano]
55500		J. I. Ruiz-Omeñaca and J. I. Canudo. 2003. Dinosaurios (Saurischia, Ornithischia) en el Barremiense (Cretácico Inferior) de la península Ibérica [Dinosaurs (Saurischia, Ornithischia) in the Barremian (Lower Cretaceous) of the Iberian peninsula]. In F. Pérez Lorente (ed.), Dinosaurios y Otros Reptiles Mesozóicos de España 269-312 [M. Carrano/M. Carrano]
16810	ETE	J. I. Ruiz-Omeñaca and J. I. Canudo. 2005. "Pleurocoelus" valdensis Lydekker 1889 (Saurischia, Sauropoda) en el Cretácico Inferior (Barremiense) de la Península Ibérica ["Pleurocoelus" valdensis Lydekker 1889 (Saurischia, Sauropoda) in the Lower Cretaceous (Barremian) of the Iberian Peninsula]. Geogaceta 38:43-45 [M. Carrano/M. Carrano]
59838		J. P. Tennant, P. D. Mannion, and P. Upchurch. 2016. Evolutionary relationships and systematics of Atoposauridae (Crocodylomorpha: Neosuchia): implications for the rise of Eusuchia. Zoological Journal of the Linnean Society 177:854-936 [P. Mannion/P. Mannion]
55315		B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]
79238		U. Villanueva-Amadoz, L. M. Sender, R. Royo-Torres, F. J. Verdú, D. Pons, L. Alcalá, and J. B. Diez. 2015. Palaeobotanical remains associated with dinosaur fossils from the Camarillas Formation (Barremian) of Galve (Teruel, Spain). Historical Biology 27(3–4):374-388 [P. Mannion/G. Varnham]