Vallipón, Castellote: Early Barremian, Spain
collected 1995

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda
Ruiz-Omeñaca et al. 2003
Bivalvia
Ruiz-Omeñaca et al. 2003
Chondrichthyes
Chondrichthyes - Hybodontiformes
Lissodus sp. Brough 1935
Ruiz-Omeñaca and Canudo 2001
Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Hybodontiformes - Polyacrodontidae
Ruiz-Omeñaca and Canudo 2001
Ruiz-Omeñaca and Canudo 2001
Polyacrodus brevicostatus (Patterson 1966)
Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Hybodontiformes - Lonchidiidae
Lissodus aff. crenulatus (Patterson 1966)
Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Synechodontiformes - Orthacodontidae
Orthacodontidae indet. de Beaumont 1960
Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Lamniformes - Eoptolamnidae
Ruiz-Omeñaca and Canudo 2001 50 specimens
Kriwet et al. 2008
Chondrichthyes - Lamniformes - Odontaspididae
Odontaspididae indet. Müller and Henle 1839
Ruiz-Omeñaca and Canudo 2001
Chondrichthyes - Rajiformes
Ruiz-Omeñaca and Canudo 2001
misspelled "mcnultyui"
Chondrichthyes - Rajiformes - Rhinobatidae
Rhinobatos cf. halteri Biddle and Landemaine 1988
Ruiz-Omeñaca and Canudo 2001
Actinopteri - Pycnodontiformes - Pycnodontidae
Ruiz-Omeñaca and Canudo 2001
Actinopteri
    = Holostei indet. Müller 1846
Ruiz-Omeñaca and Canudo 2001
Actinopteri - Lepisosteiformes
Ruiz-Omeñaca and Canudo 2001
Actinopteri - Amiiformes - Amiidae
Ruiz-Omeñaca and Canudo 2001
Amphibia - Temnospondyli
Ruiz-Omeñaca and Canudo 2001
Reptilia - Plesiosauria
Plesiosauria indet. de Blainville 1835
Ruiz-Omeñaca and Canudo 2001
Reptilia - Crocodylia
Crocodylia indet. (Owen 1842)
Reptilia - Goniopholididae
Ruiz-Omeñaca and Canudo 2001
Reptilia
Ruiz-Omeñaca and Canudo 2001
Ruiz-Omeñaca and Canudo 2001
    = aff. Theriosuchus sp. Owen 1878
Tennant et al. 2016
Reptilia - Dinosauria
Reptilia
Barco et al. 1999 1 specimen
MPZ 99/672
Ruiz-Omeñaca and Canudo 2001
Ruiz-Omeñaca and Canudo 2005
Ruiz-Omeñaca and Canudo 2001
Ruiz-Omeñaca and Canudo 2001
3 specimens
MPZ 96/50–52
Gasca et al. 2008
morphotype B
Ruiz-Omeñaca and Canudo 2001
Gasca et al. 2008
morphotype B
Reptilia - Hypsilophodontidae
2 specimens
MPZ 96/57
Reptilia
Ruiz-Omeñaca and Canudo 2001
Reptilia - Theropoda
Ruiz-Omeñaca and Canudo 2001
Reptilia
Ruiz-Omeñaca and Canudo 2001
Reptilia - Dromaeosauridae
Canudo et al. 1997 1 specimen
MPZ 96/100
Reptilia - Spinosauridae
Ruiz-Omeñaca and Canudo 2001
Reptilia - Pterosauria
Ruiz-Omeñaca et al. 1998 21 specimens
MPZ 96/59, 98/39 to 98/48; 21 teeth
Ruiz-Omeñaca et al. 1998 1 specimen
MPZ 96/58
Reptilia - Paramacellodidae
Ruiz-Omeñaca and Canudo 2001
Reptilia - Testudines
Ruiz-Omeñaca and Canudo 2001
Reptilia - Testudines - Dortokidae
Ruiz-Omeñaca and Canudo 2001
Lapparent de Broin et al. 2004
V 9326 (right posterior pleural)
Cadena and Joyce 2015
p.17
Mammalia - Multituberculata - Pinheirodontidae
Cuenca-Bescós et al. 1996 2 specimens
MPZ 96/165, 96/175
Ruiz-Omeñaca and Canudo 2001
'Some isolated teeth (I2, anterior upper premolars, and upper molars) figured and briefly described by Cuenca-Bescós et al. (1995) from the Barremian of Spain and assigned to Eobaatarinae indet. and Paulchoffatinae indet. might also belong to Lavocatia'
Mammalia - Multituberculata - Eobaataridae
Cuenca-Bescós et al. 1996 8 specimens
MPZ 96/166–173
Cuenca-Bescós et al. 1996 9 specimens
MPZ 96/156–159, MPZ2010/859–862; Three P1/3s... a ?P5 of Eobaatar may also belong to this species... other teeth such as the three upper incisors and upper and lower first and second molars may also belong to this species of Eobaatar
Badiola et al. 2012
Cuenca-Bescós et al. 1996 7 specimens
MPZ 96/160–164
    = Plagiaulacoidea indet. Ameghino 1889
Ruiz-Omeñaca and Canudo 2001
Badiola et al. 2012
MPZ 2010/863, 2010/865; Seven P1/3s resemble those of Eobaatar? pajronensis
Mammalia - Multituberculata - Paulchoffatiidae
Cuenca-Bescós et al. 1996 2 specimens
MPZ 96/174, 96/176
Mammalia - Symmetrodonta
Symmetrodonta indet. Simpson 1925
Cuenca-Bescós et al. 1996 1 specimen
MPZ 96/180
Mammalia
Cuenca-Bescós et al. 1996 3 specimens
MPZ 96/177–179
Mammalia - Eutriconodonta - Gobiconodontidae
Cuenca-Bescós and Canudo 2003
    = Gobiconodon sp. Trofimov 1978
Ruiz-Omeñaca and Canudo 2001
"sp. nov"
Docodonta
Docodonta indet. Kretzoi 1946
Cuenca-Bescós and Canudo 2003
see common names

Geography
Country:Spain State/province:Aragón County:Teruel
Coordinates: 40.8° North, 0.3° West (view map)
Paleocoordinates:26.9° North, 22.9° East (Wright 2013)
Basis of coordinate:estimated from map
Altitude:559 meters
Geographic resolution:local area
Time
Period: Cretaceous Epoch: Early Cretaceous
Stage: Barremian 10 m.y. bin: Cretaceous 2
*Period:Early/Lower Cretaceous *Epoch:Early/Lower Gallic
*International age/stage:Late/Upper Barremian
Key time interval: Early Barremian
Age range of interval: 125.77 - 121.4 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Artoles
Stratigraphic resolution:member
Stratigraphy comments: base of this formation (original name translates as "Artoles Marls and Limestones")

Villanueva-Amadoz et al. 2015: The presence of cf. Arecipites microfoveolatus Ibrahim, Afropollis sp. and Stellatopollis barghoornii Doyle from the basal levels of Artoles Formation suggests a late Barremian age (Pons et al. 2013). The schizacean fern spore content supports a Barremian age.

Bover-Arnal et al. 2016: Sr values from samples suggest a middle/late Early Barremian age for the lower-middle part of the Artoles Formation, and a Late Barremian age for the upper part. Previously the Barremian–Aptian boundary has been placed within the Artoles Formation, but late Barremian ammonites have been found in the overlying Morella Formation, ruling out an early Aptian age for the Artoles Formation.
Lithology and environment
Primary lithology: lithified calcareous sandstone
Secondary lithology: conglomerate
Includes fossils?Y
Includes fossils?Y
Lithology description: Quartitic sandstones and conglomerates. "The rock is dissolved with 10% formic acid" before screenwashing.
Environment:estuary/bay
Geology comments: "deposited in an intertidal or subtidal environment very close [related[ to the coastline"
Taphonomy
Modes of preservation:body,trace,coprolite
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:variable
Articulated whole bodies:none
Associated major elements:none
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:frequent
Bioerosion:frequent
Feeding/predation traces:gastric dissolution
Temporal resolution:time-averaged
Collection methods and comments
Collection excludes:some genera,species names
Collection methods:bulk,chemical,sieve,field collection
Minimum sieve size:0.500
Reason for describing collection:taxonomic analysis
Collection dates:1995
Collection method comments: Materials include bones, coprolites, and gastroliths. Some preparation using formic acid.
Metadata
Database number:28196
Authorizer:M. Carrano, P. Mannion, E. Vlachos, R. Benson Enterer:M. Carrano, P. Mannion, E. Vlachos, K. Maguire, R. Benson
Modifier:G. Varnham Research group:vertebrate
Created:2003-01-28 15:00:53 Last modified:2025-02-22 15:12:02
Access level:the public Released:2003-01-28 15:00:53
Creative Commons license:CC0
Reference information

Primary reference:
7706.5% 3240J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1996. Dientes de dinosaurios ornitópodos en el Barremiense superior de Castellote y Mas de las Matas (Teruel) [Ornithopod dinosaur teeth in the Upper Barremian of Castellote and Mas de las Matas (Teruel)]. Tomo Estraordinario, 125 Aniversario de la RSEHN. Real Sociedad Española de Historia Natural, Madrid 315-319 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

28224ETE A. Badiola, J. I. Canudo, and G. Cuenca-Bescós. 2007. Los fósiles de multituberculados del Hauteriviense y Barremiense de la Península Ibérica [The multituberculate fossils from the Hauterivian and Barremian of the Iberian Peninsula]. IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Salas de los Infantes, Spain 47-48 [M. Carrano/M. Carrano]
42710 A. Badiola, J. I. Canudo, and G. Cuenca-Bescós. 2012. New Early Cretaceous multituberculate mammals from the Iberian Peninsula. In P. Godefroit (ed.), Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems 409-433 [R. Benson/R. Benson]
17666ETE J. L. Barco, J. I. Canudo, J. I. Ruiz-Omeñaca and R. Royo-Torres. 1999. Bones, teeth and tracks: about sauropod dinosaur remains from Aragón (northeastern Spain). In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 22-23 [M. Carrano/M. Carrano/M. Carrano]
80368 T. Bover-Arnal, J. A. Moreno-Bedmar, G. Frijia, E. Pascual-Cebrian, and R. Salas. 2016. Chronostratigraphy of the Barremian-Early Albian of the Maestrat Basin (E Iberian Peninsula): integrating strontium-isotope stratigraphy and ammonoid biostratigraphy. Newsletters on Stratigraphy 49(1):41-68 [P. Mannion/G. Varnham/P. Mannion]
62214 E. Cadena and W. G. Joyce. 2015. A Review of the Fossil Record of Turtles of the Clades Platychelyidae and Dortokidae. Bulletin of the Peabody Museum of Natural History 56(1):3-20 [E. Vlachos/E. Vlachos/P. Mannion]
53564 J. I. Canudo, J. L. Barco, R. Royo-Torres and J. I. Ruiz-Omeñaca. 2001. Los saurópodos (Dinosauria) del Tithónico (Jurásico Superior) y del Cretácico de Aragón [The sauropoda (Dinosauria) from the Tithonian (Upper Jurassic) and Cretaceous of Aragón]. I Jornadas Internacionales sobre Paleontología de Dinosaurios y Su Entorno Actas:309-318 [M. Carrano/M. Carrano/M. Carrano]
14676ETE J. I. Canudo, G. Cuenca-Bescós, and J. I. Ruiz-Omeñaca. 1997. Dinosaurios dromaeosaúridos (Saurischia: Theropoda) en el Barremiense Superior (Cretácico Inferior) de Castellote, Teruel [Dromaeosaurid dinosaurs (Saurischia: Theropoda) in the upper Barremian (Lower Cretaceous) of Castellote, Teruel)]. Geogaceta 22:39-42 [M. Carrano/K. Maguire/M. Carrano]
17647ETE J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
41617 M. T. Carrano, R. B. J. Benson, and S. D. Sampson. 2012. The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology 10(2):211-300 [M. Carrano/M. Carrano]
54838 G. Cuenca-Bescós, J. I. Canudo, and J. I. Ruiz-Omeñaca. 1996. Los mamíferos del Barremiense superior (Cretácico inferior) de Vallipón (Castellote, Teruel) [The mammals of the upper Barremian (Lower Cretaceous) of Vallipón (Castellote, Teruel)]. Beca del Museo de Mas de las Matas Año 1995:105-137 [M. Carrano/M. Carrano]
14443ETE G. Cuenca-Bescós and J. I. Canudo. 2003. A new gobiconodontid mammal from the Early Cretaceous of Spain and its palaeogeographic implications. Acta Palaeontologica Polonica 48(4):575-582 [M. Carrano/M. Carrano/M. Carrano]
17652ETE G. Cuenca-Bescós and J. I. Canudo. 2004. Los mamíferos del Cretácico inferior de España [Lower Cretaceous mammals from Spain]. Geo-Temas 6(5):35-38 [M. Carrano/M. Carrano]
43435 V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2012. Juvenile and adult teeth of the titanosaurian dinosaur Lirainosaurus (Sauropoda) from the Late Cretaceous of Iberia. Geobios 45:265-274 [M. Carrano/M. Carrano]
31341ETE J. M. Gasca, J. I. Canudo, and M. Moreno-Azanza. 2008. Revisión de morfotipos dentales de los iguanodontoideos del Cretácico Inferior de Teruel [Revision of dental morphotypes of the iguanodontoids from the Lower Cretaceous of Teruel]. In J I Ruiz-Omeñaca, L Piñuela and J C García-Ramos (eds), XXIV Jornadas de la Sociedad Española de Paleontología, 15-18 October 2008, Museo del Jurásico de Asturias (MUJA), Colunga, Spain, Libro de Resúmenes 127-128 [M. Carrano/M. Carrano]
14365 Z. Kielan-Jaworowska, R. L. Cifelli, and Z.-X. Luo. 2004. Mammals from the age of dinosaurs: Origins, evolution, and structure. Columbia University Press, New York 1-630 [R. Whatley/R. Whatley/R. Whatley]
28298ETE J. Kriwet, S. Klug, J. I. Canudo and G. Cuenca-Bescós. 2008. A new Early Cretaceous lamniform shark (Chondrichthyes, Neoselachii). Zoological Journal of the Linnean Society 154:278-290 [M. Carrano/M. Carrano]
44196 F. d. Lapparent de Broin, X. Murelaga Bereikua, and V. Codrea. 2004. Presence of Dortokidae (Chelonii, Pleurodira) in the earliest Tertiary of the Jibou Formation, Romania: paleobiogeographical implications. Acta Palaeontologica Romaniae 4:203-215 [P. Mannion/P. Mannion]
17663ETE R. Royo-Torres, J. L. Barco, J. I. Canudo and J. I. Ruiz-Omeñaca. 2000. Descripción de un fragmento de costilla de Dinosauria (Saurópoda), del Barremiense superior del yacimiento de Vallipón (Castellote, Teruel) [Description of a dinosaur rib fragment (Sauropoda), from the upper Barremian of the Vallipón locality (Castellote, Teruel)]. Mas de las Matas 19:121-132 [M. Carrano/M. Carrano]
29947ETE J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1998. Sobre las especies de Iguanodon (Dinosauria, Ornithischia) encontradas en el Cretácico inferior de España [On the species of Iguanodon (Dinosauria, Ornithischia) found in the Lower Cretaceous of Spain]. Geogaceta 24:275-278 [M. Carrano/M. Carrano]
51920 J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 1998. Primeros restos de reptiles voladores (Pterosauria: Pterodactyloidea) en el Barremiense superior (Cretacico inferior) de Vallipón (Castellote, Teruel) [First remains of flying reptiles (Pterosauria: Pterodactyloidea) in the upper Barremian (Lower Cretaceous) of Vallipón (Castellote, Teruel)]. Mas de las Matas 17:225-249 [M. Carrano/M. Carrano]
14802ETE J. I. Ruiz-Omeñaca and J. I. Canudo. 2001. Dos yacimientos excepcionales con vertebrados continentales del Barremiense (Cretácico Inferior) de Teruel: Vallipón y La Cantalera [Two exceptional localities with continental vertebrates in the Barremian (Lower Cretaceous) of Teruel: Vallipón and La Cantalera]. Naturaleza Aragonesa 7:8-18 [M. Carrano/M. Carrano]
55492 J. I. Ruiz-Omeñaca, J. I. Canudo, and G. Cuenca-Bescós. 2002. Los dinosaurios del Jurásico y del Cretácico de la Cordillera Ibérica aragonesa [The Jurassic and Cretaceous dinosaurs of the Aragonese Iberian Cordillera]. Actas del I Encuentro Nacional de Estudios sobre la Cordillera Ibérica (ciencias de la naturaleza) 119-130 [M. Carrano/M. Carrano]
17657ETE J. I. Ruiz-Omeñaca, J. M. Canudo, and G. Cuenca-Bescós. 2003. Vallipón, el yacimiento con vertebrados continentales del Cretácico Inferior más completo de Europe [Vallipón, the most complete locality with Lower Cretaceous continental vertebrates in Europe]. El Masino [M. Carrano/M. Carrano]
55500 J. I. Ruiz-Omeñaca and J. I. Canudo. 2003. Dinosaurios (Saurischia, Ornithischia) en el Barremiense (Cretácico Inferior) de la península Ibérica [Dinosaurs (Saurischia, Ornithischia) in the Barremian (Lower Cretaceous) of the Iberian peninsula]. In F. Pérez Lorente (ed.), Dinosaurios y Otros Reptiles Mesozóicos de España 269-312 [M. Carrano/M. Carrano]
16810ETE J. I. Ruiz-Omeñaca and J. I. Canudo. 2005. "Pleurocoelus" valdensis Lydekker 1889 (Saurischia, Sauropoda) en el Cretácico Inferior (Barremiense) de la Península Ibérica ["Pleurocoelus" valdensis Lydekker 1889 (Saurischia, Sauropoda) in the Lower Cretaceous (Barremian) of the Iberian Peninsula]. Geogaceta 38:43-45 [M. Carrano/M. Carrano]
59838 J. P. Tennant, P. D. Mannion, and P. Upchurch. 2016. Evolutionary relationships and systematics of Atoposauridae (Crocodylomorpha: Neosuchia): implications for the rise of Eusuchia. Zoological Journal of the Linnean Society 177:854-936 [P. Mannion/P. Mannion]
55315 B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]
79238 U. Villanueva-Amadoz, L. M. Sender, R. Royo-Torres, F. J. Verdú, D. Pons, L. Alcalá, and J. B. Diez. 2015. Palaeobotanical remains associated with dinosaur fossils from the Camarillas Formation (Barremian) of Galve (Teruel, Spain). Historical Biology 27(3–4):374-388 [P. Mannion/G. Varnham]