PBDB Collection

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Actinopteri - Lepisosteiformes - Lepisosteidae

	Lepisosteus sp. Agassiz 1843	Astibia et al. 1990
	= Atractosteus sp. Rafinesque 1820	Pereda-Suberbiola et al. 2000
Actinopteri - Amiiformes - Palaeolabridae

	? Palaeolabrus sp. Casier 1967	Pereda-Suberbiola et al. 2000
Actinopteri - Elopiformes - Phyllodontidae

	Phyllodontidae indet. Sauvage 1875	Pereda-Suberbiola et al. 2000
Amphibia - Temnospondyli - Albanerpetontidae

	cf. Albanerpeton sp. Estes and Hoffstetter 1976	Astibia et al. 1990
	= Albanerpetontidae indet. Fox and Naylor 1982	Duffaud and Rage 1999
	five incomplete dentaries, distal part of 3 humeri
Amphibia - Temnospondyli

	Anura indet. (Fischer von Waldheim 1813)	Duffaud and Rage 1999
	distal part of 2 humeri, 8 ilia, fragmentary maxillaries, 2 broken sphenethmoids
Amphibia - Temnospondyli - Palaeobatrachidae

	Palaeobatrachidae indet. Cope 1865	Duffaud and Rage 1999
	43 prearticulars, 1 presacral vert, 2 synsacrums, 6 distal parts of humeri
Amphibia - Temnospondyli - Discoglossidae

	Discoglossidae indet. Günther 1859	Astibia et al. 1990
	6 prearticulars, 5 presacral verts, 7 sacral verts, 1 urostyle, 7 ilia, 2 humeri
Amphibia - Temnospondyli

	Salamandridae indet. Gray 1825	Astibia et al. 1990
	may be Salamandra
	= Caudata indet. Scopoli 1777	Duffaud and Rage 1999
	1 atlantal centrum, 13 fragmentary vertebrae, 1 humerus
Reptilia

	Crocodylidae indet. Cuvier 1807	Astibia et al. 1990	52 specimens
	may be Allodaposuchus
	= aff. Allodaposuchus sp. Nopcsa 1928	Pereda-Suberbiola et al. 2000
	= Allodaposuchus precedens Nopcsa 1928	Buscalioni et al. 2001
	= Allodaposuchus sp. Nopcsa 1928	Puértolas-Pascual et al. 2016

	? Trematochampsidae indet. Buffetaut 1974	Astibia et al. 1990
	= Musturzabalsuchus buffetauti Buscalioni et al. 1997	Pereda-Suberbiola et al. 2000

	Alligatoridae indet. Gray 1844	Astibia et al. 1987
	= Acynodon iberoccitanus Buscalioni et al. 1997	Pereda-Suberbiola et al. 2000

	Ischyrochampsa sp. Vasse 1995	Pereda-Suberbiola et al. 2000

	Titanosaurinae indet. Lydekker 1893	Sanz 1986	461 specimens
	orig. 1 tooth
	= Titanosauridae indet. Lydekker 1885	Astibia et al. 1990
	= Lirainosaurus astibiae n. gen., n. sp. Sanz et al. 1999	Pereda-Suberbiola et al. 2000
	Holotype: MCNA 7458, anteriormost caudal vertebra (Sanz et al., 1999: pl. 2; Díez Díaz et al., 2013: fig. 4). Paratypes: an incomplete braincase (MCNA 7439), two isolated teeth (MCNA 7440, 7441), ten dorsal vertebrae (MCNA 7442-7450, 7456), seven caudal vertebrae (MCNA 1812, 7451-7457), one scapula (MCNA 7459), one coracoid (MCNA 7460), one sternal plate (MCNA 7461), four humeri (MCNA 7462-7465), a fragment of ilium (MCNA 7466), a fragment of a pubis (MCNA 7467), three femora (MCNA 3160, 7468, 7470), a tibia (MCNA 7471), a fibula (MCNA 7472) and two osteoderms (MCNA 14473, 14474) (figured in Sanz et al., 1999).
Reptilia - Nodosauridae

	Nodosauridae indet. Marsh 1890	Astibia et al. 1990	1 specimen

	Struthiosaurus sp. Bunzel 1870	Pereda-Suberbiola 1999
	a partial skeleton, including the synsacrum, portions of the pelvic girdle and hindlimbs, a dorsal vertebra, dorsal rib fragments and a dermal scute
	= cf. Struthiosaurus austriacus Bunzel 1871	Le Loeuff and Buffetaut 1995
Reptilia - Rhabdodontidae

	Ornithopoda indet. Marsh 1881	Astibia et al. 1987
	= Rhabdodon priscus Matheron 1869	Astibia et al. 1990
	= Rhabdodon sp. Matheron 1869	Pereda-Suberbiola and Sanz 1999
	four isolated teeth, three from the maxilla and one from the dentary, two dorsal vertebrae, four caudal vertebrae, one left humerus, left and right femora
Reptilia - Hadrosauridae

	Hadrosauridae indet. Cope 1869	Pereda-Suberbiola et al. 2000
	MCNA 10510, tooth
Reptilia

	Coelurosauria indet. Huene 1914	Canudo and Ruiz-Omeñaca 2003

	cf. Paraves informal indet. A	Isasmendi et al. 2022	7 specimens
	teeth, MCNA 14523–14529

	cf. Paraves informal indet. B	Isasmendi et al. 2022	31 specimens
	teeth, MCNA 14530–14546, 14548–14561

	cf. Paronychodon sp. Cope 1876	Isasmendi et al. 2022	6 specimens
	teeth, MCNA 14547, 14562–14565, 16755

	cf. Euronychodon sp. Antunes and Sigogneau-Russell 1991	Pereda-Suberbiola et al. 2000
Reptilia - Dromaeosauridae

	Theropoda indet. Marsh 1881	Astibia et al. 1990
	= Dromaeosauridae indet. Colbert and Russell 1969	Pereda-Suberbiola et al. 2000
	3 forms (A, B,and C)

	Dromaeosauridae indet. Colbert and Russell 1969	Isasmendi et al. 2022	4 specimens
	teeth, MCNA 14623–14626

	cf. Richardoestesia informal sp. A	Isasmendi et al. 2022	35 specimens
	teeth, MCNA 14566–14610, 14612–14622, 16768, 16779, 16780, 16786–16788, 16790, 16803, 16804, 16816–16823, 16825–16830, 16832, 16833, 16837, 16841, 16846–16849, 16851, 16852, 16857–16861

	cf. Richardoestesia informal sp. B	Isasmendi et al. 2022	53 specimens
	MCNA 16765–16767, 16769–16778, 16781–16785, 16789, 16791–16802, 16805–16807, 16814, 16815, 16824, 16831, 16834–16836, 16838–16840, 16842–16845, 16850, 16853–16856
Reptilia

	Ornithomimosauria indet. Barsbold 1976	Pereda-Suberbiola et al. 2000
	= ? Ornithomimosauria indet. Barsbold 1976	Canudo and Ruiz-Omeñaca 2003
Reptilia - Abelisauridae

	? Abelisauridae indet. Bonaparte and Novas 1985	Astibia et al. 1990

	cf. Tarascosaurus sp. Le Loeuff and Buffetaut 1991	Le Loeuff and Buffetaut 1991	2 specimens
	femora, MHNV
	= Neoceratosauria indet. Novas 1991	Pereda-Suberbiola et al. 2000
	= Abelisauridae indet. Bonaparte and Novas 1985	Isasmendi et al. 2021
	femora (missing) and MCNA 14077 (caudal vertebra)

	Arcovenator sp. Tortosa et al. 2014	Isasmendi et al. 2022	32 specimens
	teeth, MCNA 1852, 1853, 1855, 2205, 2206, 8589, 8600, 10082, 14067, 14073, 14074, 14520–14522, 16756–16764, 16808–16813, 16862–16864
Reptilia - Pterosauria - Azhdarchidae

	Azhdarchidae indet. Nessov 1984	Astibia et al. 1990	295 specimens
	= cf. Azhdarcho sp. Nessov 1981	Pereda-Suberbiola et al. 2000
Reptilia - Lacertilia

	Lacertilia informal indet. 1	Rage 1999
	numerous remains (teeth)

	Lacertilia informal indet. 2	Rage 1999
	teeth

	Lacertilia informal indet. 3	Rage 1999
	teeth

	Lacertilia indet. Owen 1842	Rage 1999
	vertebrae
Reptilia - Madtsoiidae

	Madtsoia sp. Simpson 1933	Astibia et al. 1990
	= Madtsoia laurasiae Rage 1996	Rage 1999
	8 more or less complete vertebrae, and perhaps 2 fragments

	Aniliidae indet. Fitzinger 1826	Astibia et al. 1990
	= Herensugea caristiorum Rage 1996	Rage 1999
	17 vertebrae, perhaps one poorly preserved vertebra and perhaps one caudal vertebra
Reptilia - Iguanidae

	Scincomorpha informal indet. B	Astibia et al. 1990
	= Iguanidae indet. Oppel 1811	Rage 1999
	one fragment of maxilla with four teeth, two fragments of unidentified bones with two teeth, and two fragments of unidentified bone with one tooth
Reptilia

	Scincomorpha informal indet. A	Astibia et al. 1990
	= Scincomorpha indet. Camp 1923	Rage 1999
	two fragments of unidentified bones with two teeth, and one isolated tooth

	Amphisbaenia indet. Gray 1844	Astibia et al. 1990

	? Amphisbaenia indet. Gray 1844	Rage 1999
	10 vertebrae
Reptilia - Testudines

	Dortoka vasconica n. gen., n. sp. Lapparent de Broin and Murelaga 1996	Lapparent de Broin and Murelaga 1996

	? Elochelys sp. Nopsca 1931	Lapparent de Broin and Murelaga 1999
	right partial epiplastron

	Polysternon atlanticum n. sp. Lapparent de Broin and Murelaga 1996	Lapparent de Broin and Murelaga 1996	1 specimen
Reptilia

	Helochelydra nopcsai Lapparent de Broin and Murelaga 1999	Lapparent de Broin and Murelaga 1999
	partial shell, lots of fragments

	Cryptodira indet. Cope 1868	Astibia et al. 1987
	= Dermatemydidae indet. Baur 1888	Astibia et al. 1990
	"Dermatemyididae" sic
	= Solemys vermiculata Lapparent de Broin and Murelaga 1996	Joyce 2017

	Solemys vermiculata n. sp. Lapparent de Broin and Murelaga 1996	Lapparent de Broin and Murelaga 1996	260 specimens
	hypodigme around 260 shell fragments, most of them are peripherals and pleural fragments, around 25 fragments of plastron including the hologype
Mammalia

	Theria informal indet. B	Astibia et al. 1990

	Eutheria indet. (Huxley 1880)	Gheerbrant and Astibia 1999
Mammalia - Zhelestidae

	Lainodon informal new species	Gheerbrant and Astibia 2012
	L1AT 18 (talonid of left M/3) and L1AT 13 and (talonid of right M/1 or M/2)

	Theria informal indet. A	Astibia et al. 1990
	may be Marsupialia
	= Lainodon sp. Gheerbrant and Astibia 1994	Gheerbrant and Astibia 1999
	= Lainodon ragei n. sp. Gheerbrant and Astibia 2012	Gheerbrant and Astibia 2012
	L1AT 15 - type (right dP/5); referred specimens: L1AT 5: left dP/5; L1AT 15: right dP/5; L1AT 16: left dP5/; L1AT 8, left M/1 (trigonid); L1AT 9, damaged right upper molar (protocone)

	Lainodon orueetxebarriai n. gen., n. sp. Gheerbrant and Astibia 1994	Gheerbrant and Astibia 1994

	? Labes sp. Pol et al. 1992	Gheerbrant and Astibia 1999
	new species
	= Lainodon orueetxebarriai Gheerbrant and Astibia 1994	Gheerbrant and Astibia 2012
	L1AT 17, talonid of left M/1 or M/2 (fig. 2); L1AT 4, disto-labial fragment of left upper molar; and possibly L1AT11, damaged right M/1 or M/2

	Lainodon cf. orueetxebarriai Gheerbrant and Astibia 1994	Gheerbrant and Astibia 2012
	L1AT 20 (distal part of right lower premolar)
see common names

Geography

Country:	Spain	State/province:	Castilla y Leon	County:	Burgos
Coordinates:	42.7° North, 2.6° West (view map)
Paleocoordinates:	31.3° North, 8.6° East (Wright 2013)
Basis of coordinate:	estimated from map
Altitude:	747 meters
Geographic resolution:	outcrop

Time

Period:	Cretaceous	Epoch:	Late Cretaceous
Stage:	Campanian	10 m.y. bin:	Cretaceous 7
*Period:	Late/Upper Cretaceous	*Epoch:	Early/Lower Maastrichtian
Key time interval:	Late Campanian
Age range of interval:	83.6 - 72.2 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Formation:	Vitoria
Local section:	Lano section	Local bed:	S1U3
Local order:	bottom to top
Stratigraphic resolution:	bed
Stratigraphy comments: in a sector of the south limb of the Miranda-Trevino syncline, level is also called L1A lower pat of subchron C32n Also assigned to Sedano Fm. or its equivalent

Lithology and environment

Primary lithology:	"cross stratification",hardground conglomeratic sandstone
Secondary lithology:	black siltstone
Includes fossils?	Y
Includes fossils?	Y
Lithology description: Multidirectional cross-laminated sands with microconglomerates in the bottom-sets, rarely more than 1m thick. Strata become massive toward the top, and are frequently intersected by abrasion surfaces showing an iron-mineralized crust or hard-ground. The most important bone accumulation is between two of these hardgrounds. The unit is topped by two black silt beds.
Environment:	coastal indet.
Glacial or sequence phase:	regressive
Geology comments: upper foreshore and backshore environments to a subcontinental, plant-rich perhaps swampy environment. The vertebrate bearing levels indicate a continental depositional environment affected by high-energy episodes, a fluviatile environment.

Taphonomy

Modes of preservation:	body,permineralized,replaced with pyrite
Size of fossils:	macrofossils,mesofossils,microfossils
Disassociated minor elements:	all

Collection methods and comments

Collection excludes:	difficult macrofossils
Collection methods:	sieve,field collection
Collection size:	33 specimens
Reason for describing collection:	taxonomic analysis
Collection method comments: mostly chelonian shell fragments (30)

Metadata

Also known as:

Laño Quarry

Database number:

28406

Authorizer:

J. Alroy, P. Mannion, M. Carrano

Enterer:

E. Leckey, M. Carrano, P. Mannion

Modifier:

M. Carrano

Research group:

vertebrate

Collections that are a subset of this one:

37199

Created:

2003-02-12 10:11:45

Last modified:

2025-02-22 15:12:02

Access level:

the public

Released:

2003-02-12 10:11:45

Creative Commons license:

CC0

Reference information

Primary reference:

55315.

B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]

Secondary references:

7903		H. Astibia, E. Buffetaut, A. D. Buscalioni, H. Cappetta, C. Corral, R. Estes, F. Garcia-Garmilla, J. J. Jaeger, E. Jiminez-Fuentes, J. Le Loeuff, J. M. Mazin, X. Orue-Etexebarria, J. Pereda-Suberbiola, J. E. Powell, J. C. Rage, J. Rodriguez-Lazaro, J. L. Sanz and H. Tong. 1990. The fossil vertebrates from the Lano (Basque Country, Spain); new evidence on the composition and affinities of the Late Cretaceous continental faunas of Europe. Terra Nova 2:460-466 [J. Alroy/E. Leckey/M. Uhen]
7885		H. Astibia, F. Garcia-Garmilla, X. Orue-Etxebarria and J. Rodriguez-Lazaro. 1987. The Cretaceous-Tertiary boundary in a sector of the south limb of the Miranda-Trevino synclinal: the first appearance of Chelonia and Archosauria in the Basque Country. Cretaceous Research 8:15-27 [J. Alroy/E. Leckey/E. Leckey]
83613		M. Aurell, D. Torromé, J. M. Gasca, P. Calvín, M. Pérez-Pueyo, J. Parrilla-Bel, E. Medrano-Aguado, C. Martín-Closas, A. Vicente, P. Sierra-Campos, and J. I. Canudo. 2022. Latest Cretaceous palaeogeographic evolution of northeast Iberia: Insights from the Campanian continental Montalbán subbasin (Spain). Earth-Science Reviews 235:10425 [M. Carrano/M. Carrano/P. Mannion]
86530		G. Botfalvai, E. Prondvai, and A. 2021. Living alone or moving in herds? A holistic approach highlights complexity in the social lifestyle of Cretaceous ankylosaurs. Cretaceous Research 118:104633 [M. Carrano/M. Carrano]
14444	ETE	A. D. Buscalioni, F. Ortega, D. B. Weishampel and C. M. Jianu. 2001. A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia. Journal of Vertebrate Paleontology 21(1):74-86 [M. Carrano/M. Carrano/M. Carrano]
17647	ETE	J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
32749	ETE	J. Company, X. Pereda Suberbiola, and J. I. Ruiz-Omenaca. 2009. Nuevos restos fosiles del dinosaurio Lirainosaurus (Sauropoda, Titanosauria) en el Cretácico Superior (Campaniano-Maastrichtiano) de la Península Ibérica. Ameghiniana 46(2):391-405 [M. Carrano/M. Carrano/F. Aspromonte]
19147	ETE	F. M. Dalla Vecchia. 2006. Telmatosaurus and the other hadrosaurids of the Cretaceous European Archipelago. An overview. Natura Nascosta 32:1-55 [M. Carrano/M. Carrano]
56054		F. M. Dalla Vecchia. 2014. An overview of the latest Cretaceous hadrosauroid record in Europe. In D. A. Eberth & D. C. Evans (ed.), Hadrosaurs 268-297 [M. Carrano/M. Carrano]
38464		V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2011. Braincase anatomy of the titanosaurian sauropod Lirainosaurus astibiae from the Late Cretaceous of the Iberian Peninsula. Acta Palaeontologica Polonica 56(3):521-533 [M. Carrano/M. Carrano]
43435		V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2012. Juvenile and adult teeth of the titanosaurian dinosaur Lirainosaurus (Sauropoda) from the Late Cretaceous of Iberia. Geobios 45:265-274 [M. Carrano/M. Carrano]
46308		V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2013. The axial skeleton of the titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from the latest Cretaceous of Spain. Cretaceous Research 43:145-160 [P. Mannion/J. Tennant/J. Tennant]
48052		V. Díez Díaz, X. Pereda Suberbiola, and J. L. Sanz. 2013. Appendicular skeleton and dermal armour of the Late Cretaceous titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from Spain. Palaeontologia Electronica 16(2):19A [P. Mannion/J. Tennant/P. Mannion]
9509		S. Duffaud and J.-C. Rage. 1999. Amphibians from the Upper Cretaceous of Lano (Basque Country, Spain). Estudios del Museo de Ciencias Naturales de Alava 14(1):111-120 [J. Alroy/E. Leckey/J. Head]
76795		F. Escaso, A. Pérez-García, F. Ortega and J. L. Sanz. 2010. Ankylosaurian evidence from the Upper Cretaceous of South- Central Pyrenees (Lleida, Spain): a reappraisal. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 33 [M. Carrano/M. Carrano]
7914		E. Gheerbrant and H. Astibia. 1994. Un nouveau mammifere du Maastrichtien de Lano. Comptes Rendus de l'Academie des Sciences 318:1125-1131 [J. Alroy/E. Leckey/M. Uhen]
7791	5% 960	E. Gheerbrant and H. Astibia. 1999. The Upper Cretaceous mammals from Lano (Spanish Basque Country). Estudios del Museo de Ciencias Naturales de Alava 14(1):295-323 [J. Alroy/E. Leckey/E. Leckey]
44341		E. Gheerbrant and H. Astibia. 2012. Addition to the Late Cretaceous Laño mammal faunule (Spain) and to the knowledge of European “Zhelestidae” (Lainodontinae nov.). Bulletin de la Société Géologique de France 183(6):537-546 [P. Mannion/P. Mannion]
84728		E. Isasmendi, A. Torices, J. I. Canudo and X. Pereda-Suberbiola. 2021. Abelisaurid dinosaurs from the Upper Cretaceous Laño site (Iberian Peninsula). Ciências da Terra Procedia 1:38-41 [P. Mannion/P. Mannion/P. Mannion]
80599		E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
62124		W. G. Joyce. 2017. A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58(1):65-113 [P. Holroyd/P. Holroyd/P. Mannion]
9511		F. Lapparent de Broin and X. Murelaga. 1999. Turtles from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):135-211 [J. Alroy/E. Leckey/P. Mannion]
62218		F. d.e. Lapparent de Broin and X. Murelaga. 1996. Une nouvelle faune de chéloniens dans le Crétacé supérieur européen. Comptes Rendus de l’Académie des Sciences de Paris 323:729-735 [E. Vlachos/E. Vlachos]
13022	ETE	J. Le Loeuff. 1993. European titanosaurids. Revue de Paléobiologie, Volume Spéciale 7:105-117 [M. Carrano/M. Carrano/M. Carrano]
36728	ETE	J. Le Loeuff. 1998. Les dinosaures du Crétacé supérieur de l'Île Ibéro-Armoricaine [The dinosaurs of the Upper Cretaceous of the Ibero-Armorican Island]. In A. M. G. de Carvalho, A. M. Andrade, V. F. dos Santos, J. Cascalho, R. Taborda, (eds.), I Encontro Internacional sobre Paleobiologia dos Dinossáurios. Museu Nacional de História Natural, Lisboa 49-64 [M. Carrano/M. Carrano]
7842		J. Le Loeuff and E. Buffetaut. 1991. Tarascosaurus salluvicus nov. gen., nov. sp., dinosaure théropode du Crétacé supérieur du sud de la France [Tarascosaurus salluvicus nov. gen., nov. sp., a theropod dinosaur from the Upper Cretaceous of southern France]. Géobios 25(5):585-594 [M. Carrano/M. Carrano/M. Carrano]
67934		J. Le Loeuff and E. Buffetaut. 1995. The evolution of Late Cretaceous non-marine vertebrate fauna in Europe. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota, Short Papers 181-184 [M. Carrano/M. Carrano]
90698		E. Malafaia, F. Escaso, R. A. Coria, A. Pérez-García, and F. Ortega. 2025. Theropod teeth from the Upper Cretaceous of central Spain: Assessing the paleobiogeographic history of European abelisaurids. Cretaceous Research 168(78):106072 [M. Carrano/M. Carrano]
46669		A. Ösi and E. Prondvai. 2013. Sympatry of two ankylosaurs (Hungarosaurus and cf. Struthiosaurus) in the Santonian of Hungary. Cretaceous Research 44:58-63 [P. Mannion/J. Tennant/M. Carrano]
9513		X. Pereda-Suberbiola. 1999. Ankylosaurian dinosaur remains from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):273-288 [J. Alroy/E. Leckey/E. Leckey]
9559		X. Pereda-Suberbiola, H. Asibia, X. Murelaga, J. J. Elzorza, and J. J. Gomez-Alday. 2000. Taphonomy of the Late Cretaceous dinosaur-bearing beds of the Lano Quarry (Iberian Peninsula). Palaeogeography, Palaeoclimatology, Palaeoecology 157:247-275 [J. Alroy/E. Leckey/E. Leckey]
14565	ETE	X. Pereda-Suberbiola, H. Astibia, and E. Buffetaut. 1995. New remains of the armoured dinosaur Struthiosaurus from the Late Cretaceous of the Iberian peninsula (Laño locality, Basque-Cantabric basin). Bulletin de la Société Géologique de France 166(2):207-211 [M. Carrano/M. Carrano/M. Carrano]
9512		X. Pereda-Suberbiola and J. Sanz. 1999. The ornithopod dinosaur Rhabdodon from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14(1):257-272 [J. Alroy/E. Leckey/E. Leckey]
41499		A. Pérez-García, T. M. Scheyer, and X. Murelaga. 2012. New interpretations of Dortoka vasconica Lapparent de Broin and Murelaga, a freshwater turtle with an unusual carapace. Cretaceous Research 36:151-161 [R. Benson/R. Benson]
76284		E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
9510		J. Rage. 1999. Squamates (Reptilia) from the Upper Cretaceous of Lano (Basque Country, Spain). Estudios del Museo de Ciencias Naturales de Alava 14(1):121-133 [J. Alroy/E. Leckey/E. Leckey]
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70303		J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano]
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