Póvoa de Santarém: MN 6, Portugal

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Plantae indet. Haeckel 1866
c rank
Pollen (diverse)
Plantae indet. Haeckel 1866
a rank
stems and wood
Angiospermae - Ranunculales - Ranunculaceae
cf. Ranunculus sp. Linnaeus 1753
r rank
Angiospermae - Zingiberales - Zingiberaceae
Spirematospermum wetzleri Heer 1925
c rank
Authority is "(Heer)".
Angiospermae - Alismatales - Hydrocharitaceae
Stratiotes kaltennordheimensis
a rank
Authority is "(Zenker)".
Angiospermae - Typhales - Sparganiaceae
Sparganium sp. Linnaeus 1753
r rank
Angiospermae - Sapindales - Rutaceae
Toddalia maii
c rank
Authority is Gregor.
Angiospermae - Fagales - Myricaceae
Myrica cericiformis
c rank
Authority is Kownas.
Angiospermae - Vitales - Vitaceae
Vitis sp. Linnaeus 1753
r rank
Angiospermae - Nymphaeales - Nymphaeaceae
cf. Nuphar sp. Smith 1809
r rank
Pinopsida - Pinales - Pinaceae
? Picea sp. Dietrich 1824
r rank
Charales - Characeae
Characeae sp. Agardh 1824
c rank
unclassified
Metazoa indet. Haeckel 1874
r rank
coprolites
Actinopteri - Cypriniformes - Cyprinidae
Cyprinidae indet. Rafinesque 1815
r rank
Leuciscus antunesi n. sp.
Gaudant 1977 c rank
Reptilia - Squamata
Squamata indet. Oppel 1811
r rank
Reptilia - Testudines - Trionychidae
r rank
Mammalia - Cervidae
Cervidae indet. Gray 1821
r rank
Mammalia - Perissodactyla - Rhinocerotidae
Rhinocerotidae indet. Gray 1821
a rank
Mammalia - Perissodactyla - Equidae
Anchitherium aurelianense Cuvier 1825
r rank
Mammalia - Erinaceidae
Galerix cf. socialis von Meyer 1865
c rank
Mammalia - Soricidae
Miosorex grivensis Depéret 1892
r rank
Mammalia - Lagomorpha - Ochotonidae
Lagopsis verus Hensel 1856
a rank
Prolagus oeningensis (König 1825)
c rank
Mammalia - Rodentia - Sciuridae
Heteroxerus grivensis Major 1893
c rank
Miopetaurista gaillardi Mein 1970
r rank
Spermophilinus bredai (von Meyer 1848)
r rank
Mammalia - Rodentia - Cricetidae
Cricetodon sp. Lartet 1851
r rank
Cricetodon jotae Mein and Freudenthal 1971
c rank
Fahlbuschia darocensis Freudenthal 1963
c rank
Megacricetodon crusafonti Freudenthal 1963
a rank
Mammalia - Rodentia - Gliridae
Glirudinus sp. de Bruijn 1966
r rank
Peridyromys hamadryas Major 1899
r rank
Paraglis astaracensis Baudelot 1970
r rank
Pseudodryomys sp. de Bruijn 1966
r rank
Mammalia - Rodentia - Castoridae
Euroxenomys minutus von Meyer 1838
r rank
Mammalia - Proboscidea - Gomphotheriidae
Gomphotheriidae indet. Hay 1922
r rank
"Mastodonte indet."
Mammalia - Chiroptera - Vespertilionidae
Eptesicus campanensis Baudelot 1970
r rank
Bivalvia - Unionida - Unionidae
cf. Unio ribeiroi
r rank
Authority is Fontannes.
Bivalvia - Cardiida - Sphaeriidae
Pisidium sp. Pfeiffer 1821
c rank
Gastropoda - Neritidae
Theodoxus sp. Monfort 1810
r rank
Gastropoda - Bithyniidae
Bithynia sp. Leach 1818
a rank
(opercules)
Gastropoda - Heterostropha - Planorbidae
Planorbis sp. Muller 1774
c rank
(apex)
Gastropoda - Heterostropha - Testacellidae
Testacella sp. Draparnaud 1801
r rank
Gastropoda - Heterostropha - Limacidae
Limax sp. Linnaeus 1758
c rank
(limacelles)
unclassified
Limnaea sp. Lamarck 1799
c rank
(apex)
see common names

Geography
Country:Portugal State/province:Santarém County:Santarém
Coordinates: 39.3° North, 8.7° West (view map)
Paleocoordinates:39.4° North, 9.8° West
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period: Neogene Epoch: Miocene
Stage: Serravallian 10 m.y. bin: Cenozoic 5
Key time interval: MN 6
Age range of interval: 13.82 - 12.8 m.y. ago
Stratigraphy
Stratigraphic resolution:bed
Stratigraphy comments: The fossiliferous layer comes from approximately 10 meters below the surface. The presence of Anchitherium and absence of Hipparion excludes an Upper Miocene age. "..Megacricetodon crusafonti, Fahlbuschia darocensis, Cricetodon jotae are characteristic of mammalian MN6 unit, thus their ageis is nearly that of Sansan and Manchones (however the presence of Peridyromys hamadryas and Lagopsis verus do suggest, amidst this biozone, a somewhat later age than Sansan's).."

Indirect correlation with marine formations, underlying oyster-bearing beds most probably in correspondance to the apogee of the same transgression that deposed near Lisbon "schlier" facies from VI-a division division (Serravalian, Blow's zones 10-13, Globorotalia meyeri zone).
Lithology and environment
Primary lithology: lignite
Secondary lithology: mudstone
Includes fossils?Y
Includes fossils?Y
Lithology description: lignitic mudstone
Environment:pond
Glacial or sequence phase:transgressive
Geology comments: Lignite suggests marsh/stagnant pond in the vicinity of a river or ?oxbow (reference is to dead arms "des bras morts). Differing biozones are cited for original habitats of specific taxa, including savannah and forest invironments.
Taphonomy
Modes of preservation:body
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils,microfossils
Preservation of anatomical detail:good
Disassociated minor elements:all
Spatial resolution:allochthonous
Collection methods and comments
Collection methods:core,sieve,field collection
Reason for describing collection:biostratigraphic analysis
Collection method comments: In the collections of the Centro de Estratigrafia e Paleobiologia das Universidades de Lisboa.

The intial report on the lignitic clays is from a report on the geology, "Servico de Fomento Mineiro" M. Valdemiro B. Pereira, M. G. Zbyszewski the bone fragments. Antunes and Mein collected additional sediment, that was subsequently "washed" to recover the fossils.
Taxonomic list comments:r = rare
c = common
a = abundant
Metadata
Also known as:Bassin du Tage, Tagus basin, Tagus' basin, Tagus'
Database number:38137
Authorizer:J. Alroy Enterer:R. Whatley
Modifier:J. Alroy Research group:vertebrate
Created:2004-04-13 13:59:15 Last modified:2007-03-02 22:34:20
Access level:the public Released:2004-04-13 13:59:15
Creative Commons license:CC BY
Reference information

Primary reference:
10239.5% 32560M. T. Antunes and P. Mein. 1977. Contributions à la paléontologie du Miocène moyen continental du Bassin du Tage, III; Mammifères - Póvoa de Santarém, Pero Filho et Chões (Secorio), Conclusions générales. [Paleontology of the continental middle Miocene of the Tagus Basin, III; Mammals - Póvoa de Santarém, Pero Filho et Chões (Secorio), General Conclusions. Ciencias de Terra (UNL) (3)143-165 [J. Alroy/R. Whatley/J. Head]

Secondary references:

10244 J. Gaudant. 1977. Contributions à la paléontologie du Miocène moyen continental du Bassin du Tage, II; Observations sur les dents pharyngiennes de Poissons Cyprinides - Póvoa de Santarém. [Paleontology of the continental middle Miocene of the Tagus Basin, II; Observations on the pharyngeal teeth of cyprinid fish - Póvoa de Santarém]. Ciencias de Terra (UNL) (3)129-141 [J. Alroy/R. Whatley/R. Whatley]