PBDB Collection

Czatkowice 1 quarry: Olenekian, Poland
collected 1978

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Osteichthyes

	Osteichthyes indet. Huxley 1880
Dipnoi - Gnathorhizidae

	Gnathorhiza sp. Cope 1883	Borsuk-Bialynicka et al. 2003
Reptilia - Procolophonidae

	Procolophonidae informal unnamed genus 1	Borsuk−Białynicka and Lubka 2009
	Maxillae ZPAL RV/734; ZPAL RV/739; dentary: ZPAL RV/733; vertebrae type B2 (Lubka 1999): ZPAL RV/840–842, 849, 851–853, 860, and numerous unnumbered specimens.

	Procolophonidae informal unnamed genus 2	Borsuk−Białynicka and Lubka 2009
	Dentary fragment ZPAL RV/507; Vertebrae type B3 (Lubka 1999): ZPAL RV/767–772; ZPAL RV/846–870, and several unnumbered vertebrae

	Procolophonidae informal unnamed genus 3	Borsuk−Białynicka and Lubka 2009	1 element
	ZPAL RV/716, dentary

	Procolophonida indet.
	= Procolina teresae n. gen., n. sp. Borsuk−Białynicka and Lubka 2009	Borsuk−Białynicka and Lubka 2009
	Many disarticulated premaxillae, maxillae, dentaries, palatal bones and quadrates, vertebrae referred to as type B1 vertebrae by Lubka (1999), and numerous bone fragments
Reptilia

	Prolacertiformes indet. Camp 1945
	= Czatkowiella harae n. gen., n. sp. Borsuk−Białynicka and Evans 2009	Borsuk−Białynicka and Evans 2009
	ZPAL R.V/100, right maxilla. Many referred specimens

	Archosauriformes indet. Gauthier 1986	Borsuk−Białynicka and Sennikov 2009

	Collilongus rarus n. gen., n. sp. Borsuk−Białynicka and Sennikov 2009	Borsuk−Białynicka and Sennikov 2009	20 elements
	Four cervicals: ZPAL RV/579, 580, 581, 596; ?four dorsals: ZPAL RV/584, 585, 588, 694; one sacral ZPAL RV/1369; and twelve caudals: ZPAL RV/583, 594, 661, 662, 663, 1362, 1363, 1364, 1365, 1366, 1367, 1368
Reptilia - Euparkeriidae

	Archosauria indet. Cope 1869		260 specimens
	= Osmolskina czatkowicensis n. gen., n. sp. Borsuk-Bialynicka and Evans 2003	Borsuk-Bialynicka and Evans 2003
	Approximately 100 cranial elements. Catalogued postcranial specimens are: 63 measured vertebrae (Supplement 1); twelve scapulae: ZPAL RV/881–885, 902, 1161, 1261, 1282–1286; eight coracoids: ZPAL RV/890–891, 903, 1166–1169, 1311; eleven humeri ZPAL RV/1164, 1262–1263, 1266–1272, 1277; ten ilia: ZPAL RV/630, 678, 679, 911–913, 918, 919, 924, 925; four ischia: ZPAL RV/887–889, 892; five pubes: 905–907, 909, 910; fifteen femora: ZPAL RV/938–940, 1252, 1254, 1264, 1287–1302, 1304; 18 tibiae: ZPAL RV/1165, 1171, 1176, 1177, 1221, 1222, 1257–1260, 1265, 1303, 1306, 1317–1321, fragmentary radius ZPAL RV/1232 and ulnae ZPAL RV/1230 and 1279; fragmentary fibulae ZPAL RV/1182, 1225, 1247; numerous tarsal bones, of which the astragalus: ZPAL RV/811, a calcaneum ZPAL RV/810, and a fourth distal tarsal ZPAL RV/812 are catalogued and figured.
Reptilia

	Lepidosauromorpha indet. Benton 1983		84 elements
	= Sophineta cracoviensis n. gen., n. sp. Evans and Borsuk−Białynicka 2009	Evans and Borsuk−Białynicka 2009
	ZPAL RV/175, a nearly complete right maxilla (holotype). ZPAL RV/226–230, 445, 1086 (frontals); ZPAL RV/231, 232, 443 (parietals); ZPAL RV/1056 (prefrontal); ZPAL RV/233–235, 966, 967, 1060 (postfrontals); ZPAL RV/236–238, 1058, 1062 (postorbitals); ZPAL RV/10, 239–244, 965, 1054, 1059 (jugals); ZPAL RV/245–249, 1055, 1057, 1069 (squamosals); ZPAL RV/968–973 (pterygoids); ZPAL RV/13, and 974 (quadrates); ZPAL RV/3, 7, 174–178, 455, 472, 493, 823, 824 (maxillae); ZPAL RV/1061 (premaxilla); ZPAL RV/174, 189, 392, 493, 512, 1101 (dentaries); ZPAL RV/23, 627, 628, 1079, 1080, 1098, 1108, 1110, 1121 (vertebrae); ZPAL RV/948–950, 952, 959, 1053, 1063 (ilia).
Reptilia - Kuehneosauridae

	Diapsida indet. Osborn 1903		114 elements
	= Pamelina polonica n. gen., n. sp. Evans 2009	Evans 2009
	ZPAL RV/1036 (maxilla, holotype). ZPAL RV/378, 381, 383–384, 387, 537, 1087 (frontals); ZPAL RV/157, 975–978, 1028 (parietals); ZPAL RV/ 979, 980, 1001, 1002 (prefrontals); ZPAL RV/148, 1003, 1004, 1006, 1007, 1027 (postfrontals); ZPAL RV/806, 1005, 1072, 1077, 1078 (postorbitals); ZPAL RV/1008–1010 (jugals); ZPAL RV/147, 366, 1011–1026 (squamosals); ZPAL RV/1029–1033, 1083 (quadrates); ZPAL RV/1, 6, 1034–1042, 1081 (maxillae); ZPAL RV/146, 151–155, 184, 201, 451, 1043–1045, 1082 (premaxillae); ZPAL RV/1048–1050 (sphenoids); ZPAL RV/142–144, 149, 162, 185, 186, 1046, 1047 (dentaries); ZPALRV/1066, 67 (possible palatines); ZPAL RV/ 613, 617, 627, 1194–1210 (vertebrae); ZPAL RV/1211–1213 (ribs); ZPAL RV/555, 981 (ilia).
Amphibia - Temnospondyli

	Czatkobatrachus polonicus n. gen., n. sp. Evans and Borsuk-Bialynicka 1998	Evans and Borsuk-Bialynicka 1998	76 elements
Amphibia - Temnospondyli - Tupilakosauridae

	Batrachosuchoides sp. Shishkin 1967	Shishkin and Sulej 2009	8 elements
	A series of incomplete isolated bones, collection ZPAL AbIV: postparietal 101; ectoptery− goids 36, 53; exoccipitals 48, 120, 152; surangular 69; clavicle 61
Amphibia - Temnospondyli

	Amphibia indet. Linnaeus 1758		53 elements
	= Parotosuchus speleus n. sp. Shishkin and Sulej 2009	Shishkin and Sulej 2009
	ZPAL AbIV/105, left vomer (holotype). A series of isolated bones and bone fragments from the type locality, all belonging to collection ZPAL AbIV. The specimens include: postfrontal 43; postorbital 151; vomers 37, 38, 58, 59, 65–67, and 73; palatine 97; ectopterygoids 46, 50, and 107; exoccipitals 33 and 104; premaxillae 62 and 92; maxillae 31,106,116,118, and 153; maxillae or dentaries (poorly identifiable fragments) 47, 51, 52, 57, 60, 64, 74, 91, and 93; dentaries 32, 34, 35, 82, and 119; angular 63; surangular 68; clavicles 75 and 87; interclavicle 102; neural arches 158–160; scapulae (?) 88 and 103; humeri 84 and 86; radius (?) 96; ribs 85 and 99.
see common names

Geography

Country:	Poland	State/province:	Malopolskie	County:	Krakowski
Coordinates:	50.1° North, 19.6° East (view map)
Paleocoordinates:	17.8° North, 26.3° East
Basis of coordinate:	based on nearby landmark
Altitude:	318 meters
Geographic resolution:	outcrop

Time

Period:	Triassic	Epoch:	Early Triassic
Stage:	Olenekian	10 m.y. bin:	Triassic 1
Key time interval:	Olenekian
Age range of interval:	251.2 - 247.2 m.y. ago

Stratigraphy

Stratigraphic resolution:	bed
Stratigraphy comments: earliest Late Olenekian (Yarenskian) on the basis of tetrapod (temnospondyl) biostratigraphy (Shishkin and Sulej 2009)

Lithology and environment

Primary lithology:	fine,yellow sandy "limestone"
Secondary lithology:	"limestone"
Includes fossils?	Y
Lithology description: "fine grained sandy yellow limestone with some calcite cement [with] bones, occasional clasts, and concretions. A few samples show evidence of sedimentary bedding and parallel orientation of the bones, but this is not generally the case." bone breccia within a Carboniferous limestone
Environment:	fissure fill
Geology comments: single karst fissure; arid environment with sparse vegetation and occasional pools with aquatic communities

Taphonomy

Modes of preservation:	body
Lagerst�tten type:	concentrate
Degree of concentration:	-multiple events
Size of fossils:	macrofossils,mesofossils
Spatial orientation:	random
Preservation of anatomical detail:	excellent
Articulated whole bodies:	none
Associated major elements:	none
Disassociated major elements:	many
Disassociated minor elements:	many
Size sorting:	medium
Fragmentation:	frequent
Temporal resolution:	time-averaged
Spatial resolution:	allochthonous

Collection methods and comments

Collection methods:	selective quarrying,chemical,mechanical,field collection
Reason for describing collection:	general faunal/floral analysis
Collection dates:	1978
Collection method comments: ZPAL: Institute of Paleobiology, Warsaw

Metadata

Database number:	38528
Authorizer:	M. Carrano, R. Butler	Enterer:	M. Carrano, R. Butler
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2004-04-23 10:17:39	Last modified:	2021-10-19 10:57:09
Access level:	the public	Released:	2004-04-23 10:17:39
Creative Commons license:	CC BY

Reference information

Primary reference:

10334.

M. Borsuk-Bialynicka, E. Cook, S. E. Evans and T. Maryanska. 1999. A microvertebrate assemblage from the Early Triassic of Poland. Acta Palaeontologica Polonica 44(2):167-188 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

38012		M. Borsuk−Białynicka and S. E. Evans. 2009. A long−necked archosauromorph from the Early Triassic of Poland. Palaeontologia Polonica 65:203-234 [R. Butler/R. Butler]
38013		M. Borsuk−Białynicka and S. E. Evans. 2009. Cranial and mandibular osteology of the Early Triassic archosauriform Osmolskina czatkowicensis from Poland. Palaeontologia Polonica 65:235-281 [R. Butler/R. Butler]
38010		M. Borsuk−Białynicka and M. Lubka. 2009. Procolophonids from the Early Triassic of Poland. Palaeontologia Polonica 65:107-144 [R. Butler/R. Butler]
38014		M. Borsuk−Białynicka and A. G. Sennikov. 2009. Archosauriform postcranial remains from the Early Triassic karst deposits of southern Poland. Palaeontologia Polonica 65:283-328 [R. Butler/R. Butler]
10333		M. Borsuk-Bialynicka and S. E. Evans. 2003. A basal archosauriform from the Early Triassic of Poland. Acta Palaeontologica Polonica 48(4):649-652 [M. Carrano/M. Carrano/M. Carrano]
10336		M. Borsuk-Bialynicka, T. Maryanska, and M. A. Shishkin. 2003. New data on the age of the bone breccia from the locality Czatkowice 1 (Cracow Upland, Poland). Acta Palaeontologica Polonica 48(1):153-155 [M. Carrano/M. Carrano/M. Carrano]
38007		E. Cook and C. Trueman. 2009. Taphonomy and geochemistry of a vertebrate microremains assemblage from the Early Triassic karst deposits at Czatkowice 1, southern Poland. Palaeontologia Polonica 65:17-30 [R. Butler/R. Butler]
38011		S. E. Evans. 2009. An early kuehneosaurid reptile (Reptilia: Diapsida) from the Early Triassic of Poland. Palaeontologia Polonica 65:145-178 [R. Butler/R. Butler]
35304		S. E. Evans and M. Borsuk−Białynicka. 2009. A small lepidosauromorph reptile from the Early Triassic of Poland. Palaeontologia Polonica 65:179-202 [R. Butler/R. Butler]
38009		S. E. Evans and M. Borsuk−Białynicka. 2009. The Early Triassic stem−frog Czatkobatrachus from Poland. Palaeontologica Polonica 65:79-105 [R. Butler/R. Butler]
10335		S. E. Evans and M. Borsuk-Bialynicka. 1998. A stem-group frog from the Early Triassic of Poland. Acta Palaeontologica Polonica 43(4):573-580 [M. Carrano/M. Carrano/M. Carrano]
78367		M. Paszkowski. 2009. The Early Triassic karst of Czatkowice 1, southern Poland. Palaeontologia Polonica 65:7-16 [M. Carrano/M. Carrano]
38008		M. A. Shishkin and T. Sulej. 2009. The Early Triassic temnospondyls of the Czatkowice 1 tetrapod assemblage. Palaeontologia Polonica 65:31-77 [R. Butler/R. Butler]